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Rhinoceros (genus)

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Genus of mammals
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Rhinoceros
Temporal range: Late Miocene-Recent 9–0 Ma
Indian rhinoceros (R. unicornis)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Rhinocerotidae
Tribe: Rhinocerotini
Genus: Rhinoceros
Linnaeus, 1758
Type species
Rhinoceros unicornis
Linnaeus, 1758
Species

Rhinoceros is a genus comprising one-horned rhinoceroses. This scientific name was proposed by Swedish taxonomist Carl Linnaeus in 1758.[1] The genus contains two species, the Indian rhinoceros (Rhinoceros unicornis) and the Javan rhinoceros (Rhinoceros sondaicus). Although both members are threatened, the Javan rhinoceros is one of the most endangered large mammals in the world with only 60 individuals surviving in Java (Indonesia). The word 'rhinoceros' means "nose-horn" in Ancient Greek.[clarification needed ]

Etymology

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The genus name Rhinoceros comes from the Ancient Greek words ῥινο- (rhino-), meaning "of the nose" and κέρας (kerás), meaning "horn".[2] [3]

Classification

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The genus Rhinoceros comprises:

The earliest remains of the genus are known from the Late Miocene, represented by remains such as an indeterminate species found in deposits in Myanmar dating to around 8-9 million years ago.[9]

The species "Rhinoceros" philippinensis from the early Middle Pleistocene of the Philippines and "Rhinoceros" sinensis hayasakai from the Early-Middle Pleistocene of Taiwan have been transferred to Nesorhinus, which appears to be closely related to Rhinoceros.[10] While Rhinoceros fusuiensis Yan et al. 2014[11] from the Early Pleistocene of South China has been transferred to Dicerorhinus.[10] [12]

 Relationships among Late Pleistocene and modern rhinoceros genera, based on nuclear DNA, after Liu et al, 2021:[13]

Elasmotheriinae

Elasmotherium sibiricum

Rhinocerotinae

Black rhinoceros (Diceros bicornis)

White Rhinoceros (Ceratotherium simum)

Rhinoceros

Indian rhinoceros (Rhinoceros unicornis)

Javan rhinoceros (Rhinoceros sondaicus)

Sumatran rhinoceros (Dicerorhinus sumatrensis)

Woolly rhinoceros (Coelodonta antiquitatis)

Merck's rhinoceros (Stephanorhinus kirchbergensis)

Bayesian morphological phylogeny after Pandolfi, 2023 Note: This excludes living African rhinoceros species.[14]

Rhinocerotina
Dicerorhinus
Coelodonta
Pliorhinus Stephanorhinus

Stephanorhinus jeanvireti

Stephanorhinus etruscus

Stephanorhinus hundsheimensis

Stephanorhinus hemitoechus (Narrow-nosed or steppe rhinoceros)

Stephanorhinus kirchbergensis (Merck's or forest rhinoceros)

References

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  1. ^ a b Linnæus, C. (1758). "Rhinoceros unicornis". Caroli Linnæi Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Holmiae: Salvius. p. 56.
  2. ^ Liddell, H. G. & Scott, R. (1940). "ῥίς". A Greek-English Lexicon (Revised and augmented ed.). Oxford: Clarendon Press.
  3. ^ Liddell, H. G. & Scott, R. (1940). "κέρᾳ". A Greek-English Lexicon (Revised and augmented ed.). Oxford: Clarendon Press.
  4. ^ Desmarest, A. G. (1822). "Rhinocéros des Îles de La Sonde". Mammalogie, ou, Description des espèces de mammifères. Vol. 2. Paris: Mme Agasse. pp. 399–400.
  5. ^ a b Pandolfi, Luca; Maiorino, Leonardo (2016年02月06日). "Reassessment of the largest Pleistocene rhinocerotine Rhinoceros platyrhinus (Mammalia, Rhinocerotidae) from the Upper Siwaliks (Siwalik Hills, India)" (PDF). Journal of Vertebrate Paleontology. 36 (2): e1071266. Bibcode:2016JVPal..36E1266P. doi:10.1080/02724634.2015.1071266. ISSN 0272-4634. S2CID 86893957.
  6. ^ Schepartz, L. A.; Miller-Antonio, S. (2010). "Taphonomy, life history, and human exploitation of Rhinoceros sinensis at the Middle Pleistocene site of Panxian Dadong, Guizhou, China". International Journal of Osteoarchaeology. 20 (3): 253–268. doi:10.1002/oa.1025.
  7. ^ Antoine, Pierre-Olivier (March 2012). "Pleistocene and Holocene rhinocerotids (Mammalia, Perissodactyla) from the Indochinese Peninsula". Comptes Rendus Palevol. 11 (2–3): 159–168. Bibcode:2012CRPal..11..159A. doi:10.1016/j.crpv.201103002.
  8. ^ Tong, Hao-wen (November 2012). "Evolution of the non-Coelodonta dicerorhine lineage in China". Comptes Rendus Palevol. 11 (8): 555–562. Bibcode:2012CRPal..11..555T. doi:10.1016/j.crpv.201206002.
  9. ^ Longuet, Morgane; Zin-Maung-Maung-Thein; Thaung-Htike; Man-Thit-Nyein; Takai, Masanaru (2024年08月02日). "New fossil remains of Rhinocerotidae (Perissodactyla) from the early Late Miocene Tebingan area, central Myanmar". Historical Biology. 36 (8): 1468–1481. doi:10.1080/08912963.2023.2218873. ISSN 0891-2963.
  10. ^ a b Antoine, P.-O.; Reyes, M. C.; Amano, N.; Bautista, A. P.; Chang, C.-H.; Claude, J.; De Vos, J.; Ingicco, T. (2021). "A new rhinoceros clade from the Pleistocene of Asia sheds light on mammal dispersals to the Philippines". Zoological Journal of the Linnean Society. 194 (2): 416–430. doi:10.1093/zoolinnean/zlab009.
  11. ^ Yan, Yaling; Wang, Yuan; Jin, Changzhu; Mead, Jim I. (December 2014). "New remains of Rhinoceros (Rhinocerotidae, Perissodactyla, Mammalia) associated with Gigantopithecus blacki from the Early Pleistocene Yanliang Cave, Fusui, South China". Quaternary International. 354: 110–121. Bibcode:2014QuInt.354..110Y. doi:10.1016/j.quaint.201401004.
  12. ^ Pandolfi, Luca (2023年01月19日). "Reassessing the phylogeny of Quaternary Eurasian Rhinocerotidae". Journal of Quaternary Science. 38 (3): 291–294. Bibcode:2023JQS....38..291P. doi:10.1002/jqs.3496. hdl:11563/163194 . ISSN 0267-8179. S2CID 256167036.
  13. ^ Liu, Shanlin; Westbury, Michael V.; Dussex, Nicolas; Mitchell, Kieren J.; Sinding, Mikkel-Holger S.; Heintzman, Peter D.; Duchêne, David A.; Kapp, Joshua D.; von Seth, Johanna; Heiniger, Holly; Sánchez-Barreiro, Fátima (16 September 2021). "Ancient and modern genomes unravel the evolutionary history of the rhinoceros family". Cell. 184 (19): 4874–4885.e16. doi:10.1016/j.cell.2021年07月03日2 . hdl:10230/48693 . PMID 34433011. S2CID 237273079.
  14. ^ Pandolfi, Luca (April 2023). "Reassessing the phylogeny of Quaternary Eurasian Rhinocerotidae". Journal of Quaternary Science. 38 (3): 291–294. Bibcode:2023JQS....38..291P. doi:10.1002/jqs.3496 . hdl:11563/163194 . ISSN 0267-8179.
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Incertae sedis
Anthracobunia
Anthracobunidae
Cambaytheriidae
Brontotheriidae
Equoidea
Equidae
Palaeotheriidae
Pachynolophinae
Palaeotheriinae
Hyopsodontidae?
Phenacodontidae?
Meniscotheriinae
Phenacodontinae
Tapiromorpha
    • See below↓
Incertae sedis
Isectolophidae
Homogalaxinae
Isectolophinae
Ancylopoda
Ceratomorpha
Tapiroidea
Deperetellidae
Helaletidae
Tapiridae
Rhinocerotoidea
    • See below↓
Rhinocerotoidea sensu lato
Indolophidae
Rhinocerotoidea
sensu stricto
Hyrachyidae
Eggysodontidae
Hyracodontidae
Amynodontidae
Amynodontinae
Cadurcodontini
Metamynodontini
Paraceratheriidae
Forstercooperiinae
Paraceratheriinae
Rhinocerotidae
    • See below↓
Aceratheriinae
Elasmotheriinae
Diceratheriini
Elasmotheriini
Rhinocerotinae
Teleoceratini
Rhinocerotina
Dicerorhinini
Dicerotini
Rhinocerotini
Extant Perissodactyla (Odd-toed ungulates) species by suborder
Equidae
(Horse family)
Equus
(including Zebras)
Rhinocerotidae
(Rhinoceroses)
Rhinoceros
Dicerorhinus
Ceratotherium
Diceros
Tapiridae (Tapirs)
Tapirus


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