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Scotokaryotes

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Theoretical clade in the Eukaryota taxonomical domain
Scotokaryotes
Scientific classification Edit this classification
Domain: Eukaryota
(unranked): Neokaryotes
(unranked): Scotokaryotes
Cavalier-Smith, 1999
Clades
Synonyms
  • Opimoda Derelle et al., 2015[1]

The Scotokaryotes is a proposed basal eukaryote clade sister to Diaphoretickes.[2] Likely Scotokaryote groupings include the Metamonads, the Malawimonas and the Podiata.[3] [4] The name comes from the Greek word scotos (dark) due to the complete absence of photosynthetic organisms in this clade, contrasting with Diaphoretickes and Excavata.[2]

An alternative to the UnikontBikont division was suggested by Derelle et al. in 2015,[1] where they proposed the acronyms Opimoda–Diphoda respectively, as substitutes to the older terms (Opimoda includes the old 'unikonts', plus some former bikonts; Diphoda includes most of the old 'bikonts', but not all). The name Opimoda is formed from the letters (shown in capitals) of OPIsthokonta and aMOebozoa.

Sarcomastigota Cavalier-Smith, 1983 is a proposed subkingdom (currently shown to be paraphyletic) that includes all members of Opimoda that are not animals or fungi.[5] Sulcozoa Cavalier-Smith, 2012 is a proposed phylum (currently shown to be paraphyletic) within Sarcomastigota that does not include the phyla Amoebozoa (clade) and Choanozoa (paraphyletic), i.e. it includes the proposed subphyla Apusozoa and Varisulca.[6]

Phylogeny

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A proposed cladogram is[7]

Eukaryota
Diphoda Opimoda


See also

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References

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  1. ^ a b Derelle, Romain; Torruella, Guifré; Klimeš, Vladimír; Brinkmann, Henner; Kim, Eunsoo; Vlček, Čestmír; Lang, B. Franz; Eliáš, Marek (17 February 2015). "Bacterial proteins pinpoint a single eukaryotic root". Proceedings of the National Academy of Sciences. 112 (7): E693–E699. Bibcode:2015PNAS..112E.693D. doi:10.1073/pnas.1420657112 . PMC 4343179 . PMID 25646484.
  2. ^ a b Cavalier-Smith, Thomas; Chao, Ema E.; Lewis, Rhodri (December 2015). "Multiple origins of Heliozoa from flagellate ancestors: New cryptist subphylum Corbihelia, superclass Corbistoma, and monophyly of Haptista, Cryptista, Hacrobia and Chromista". Molecular Phylogenetics and Evolution. 93: 357. doi:10.1016/j.ympev.201507004 .
  3. ^ Cavalier-Smith, Thomas (2013年05月01日). "Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa". European Journal of Protistology. 49 (2): 115–178. doi:10.1016/j.ejop.201206001. PMID 23085100.
  4. ^ Cavalier-Smith, Thomas (2017). "Euglenoid pellicle morphogenesis and evolution in light of comparative ultrastructure and trypanosomatid biology: semi-conservative microtubule/strip duplication, strip shaping and transformation". European Journal of Protistology. 61 (Pt A): 137–179. doi:10.1016/j.ejop.201709002 . PMID 29073503.
  5. ^ Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M. (2015年06月11日). "Correction: A Higher Level Classification of All Living Organisms". PLOS ONE. 10 (6) e0130114. Bibcode:2015PLoSO..1030114R. doi:10.1371/journal.pone.0130114 . ISSN 1932-6203. PMC 5159126 . PMID 26068874.
  6. ^ Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M. (2015年04月29日). "A Higher Level Classification of All Living Organisms". PLOS ONE. 10 (4) e0119248. Bibcode:2015PLoSO..1019248R. doi:10.1371/journal.pone.0119248 . ISSN 1932-6203. PMC 4418965 . PMID 25923521..
  7. ^ Torruella, Guifré; Galindo, Luis Javier; Moreira, David; López-García, Purificación (2025). "Phylogenomics of neglected flagellated protists supports a revised eukaryotic tree of life". Current Biology. 35 (1): 198–207.e4. doi:10.1016/j.cub.2024年10月07日5. hdl:10481/102343 . ISSN 0960-9822.
Eukaryote classification
Amoebozoa
Holomycota
Filozoa
Choanozoa
Haptista
    SAR    
Rhizaria
Alveolata
Myzozoa
Stramenopiles
Bigyra *
Gyrista
Pancryptista
Cryptista
Archaeplastida
(plants sensu lato)
Viridiplantae
(green plants or
plants sensu stricto)
Streptophyta
Provora
Membrifera
Discoba
Discicristata
Metamonada *
Malawimonadida
Ancyromonadida
CRuMs
Genera of
uncertain affiliation
Acritarchs
and other fossils

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