The USCSS Prometheus was the flagship of the Weyland Corporation fleet, being one of the first ships capable of interstellar travel. It was designed by Peter Weyland and was the ship he boarded on his fateful journey to LV-223 during the events of the Prometheus film.
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The Prometheus lands on LV-223.
The USCSS Prometheus took off from Earth some time in 2091, after archeologists Elizabeth Shaw and Charlie Holloway discover indications of an alien species called the Engineers inhabiting a far-off system. Peter Weyland secretly joined the crew in the hopes of convincing the Engineers to help him conquer death.
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The Prometheus crashes into the Engineer's ship.
When the ship arrives at LV-223, the crew discovers a mysterious Pyramid they hope will hold the secrets of the Engineers. Much to Weyland's disappointment, it turns out to be a bioweapons facility used to store Chemical A0-3959X.91-1, also known as Black Goo . Rather than grant his wish, a remaining Engineer kills Weyland and his compatriots. Shaw escapes and warns the captain of the Prometheus, Janek , about the ship. He responds by colliding the Prometheus into the Engineer's ship, destroying them both in the process.
Prometheus: Cast of Characters Here's a quick look of the characters from 2012's Prometheus. Some spoilers follow... Dr. Elizabeth Shaw (played by by Noomi Rapace)<p> Archeologist Elizabeth Shaw and her lover/partner Charlie Holloway join an intergalactic mission to LV-223 based on their research of ancient Earth star maps, and convincing CEO Peter Weyland that the maps are an "invitation" for humans to seek out their galactic creators, the Engineers. <br> Investigating a Pyramid on LV-223, Shaw and company discover that the Engineers were all wiped out by their own bioweapon. David, Weyland's Synthetic "son," sees fit to poison Holloway with Black Goo harvested from the Pyramid. Tainted with the alien substance, Holloway and Shaw make love, which results in Holloway mutating (and being killed by Weyland exec Meredith Vickers) and Shaw experiencing a rapid alien pregnancy. <br> Shaw removes her alien fetus - the Trilobite - using a surgical MedPod and then discovers that the elderly Weyland has been on board the Prometheus ship the entire time, with hopes of talking to an Engineer and convincing it to grant him immortality. Shaw accompanies Weyland, David, and others back into the Pyramid, to an underground Engineer ship, to awaken a hyper-sleeing Engineer.<br> The Engineer kills everyone (dismantling David in the process) and starts up its ship, attempting to finish what it started which was the extermination of life on Earth. Shaw escapes the Engineer's ship, narrowly escapes that ship crashing down to the planet (when Prometheus pilot Janek uses the Prometheus as a battering ram) and makes her way to Vickers' lifeboat pod. There she watches as the now-grown Trilobite defeats the Engineer (who survived his ship's crash). <br> Shaw then finds David's parts, forming an agreement with him to fly a different Engineer ship to the Engineer's origin planet to find out why they wanted to kill off humanity. David (played by Michael Fassbender)</p> David is Peter Weyland's Synthetic creation, the first of his kind, and a type of surrogate son for Weyland. While the Prometheus crew is in cryosleep, David minds the ship, practicing ancient Earth languages, spying on Dr. Elizabeth Shaw's dreams about her late father, and watching Peter O' Toole in Lawrence of Arabia. <br> David is dutiful, but ultimately is operating at the behest of the elderly Weyland, who's secretly on board the ship, hoping to meet a cosmic Engineer and discover the secret to immortality. David also holds humans in slight contempt, and gets into small passive-aggressive exchanges with Dr. Charlie Holloway. <br> David, unbeknownst to others, harvest Black Goo from the Pyramid on LV-223 and then spikes Holloway's drink with it, just to see what happens. Holloway has sex with Dr. Elizabeth Shaw soon drinking the substance and it results in Holloway being poisoned, mutated, and killed while Shaw becomes pregnant with a Trilobite fetus. David informs Shaw she's pregnant and suggests she enter cryosleep so that her "baby" can be removed back on Earth but she takes matters into her own hands and removes it via MedPod.<br> Previously, in a bit of side exploration in the Pyramid, David uncovers the Engineers' ship and found out that one Engineer was still alive in cryosleep - and also that the cosmic creators' plan of action was to fly to Earth and release the Black Goo pathogen on the populous, wiping out all life (while also mutating a portion of it). Knowing all this, he still brings the old and dying Weyland to meet the Engineer. The awakened Engineer destroys David, kills Weyland, and resumes its mission to annihilate all life on Earth.<br> After the Engineer's ship is downed and the Engineer himself is killed by the grown trilobite, David, now just a head, contacts the surviving Dr. Shaw, offering a deal to fly them both away if she recovers his body. She agrees, but says she wants to go to the Engineers' home planet and demand answers. Dr. Charlie Holloway (played by Logan Marshall-Green)<p> Archeologist Dr. Charlie Holloway and his lover/parter Elizabeth Shaw join an intergalactic mission to LV-223 based on their research of ancient Earth star maps, and convincing CEO Peter Weyland that the maps are an "invitation" for humans to seek out their galactic creators, the Engineers. <br> Charlie is thrilled to meet the Engineers, the creators of humankind, but it upset to learn, from Weyland exec Meredith Vickers, that he and Shaw are not to initiate any contact with the alien beings. Holloway becomes even more dejected when he finds all the Engineers dead inside of a mysterious Pyramid. During his time on LV-223, Holloway also trades gentle barbs with David, Peter Weyland's Synthetic "son," teasing him about being a robot. This is probably the reason why David chooses Holloway as the one to test the Black Goo on, tainting Holloways drink with a Black Goo drop while the archeologist is on a bender. <br> Not knowing he's been infected, Holloway then has sex with Shaw. The next morning, back in the Pyramid, Holloway starts feeling himself painfully mutate. They rush Holloway back to Prometheus, but Vickers refuses to let him on board due to quarantine rules. Holloway knows that he's dying, or at least changing into something else, so he sacrifices himself, walking toward Vickers, telling her to use her flamethrower on him. She obliges and torches him to death. Meredith Vickers (played by Charlize Theron)<p> Meredith Vickers is Peter Weyland's actual child, his daughter, though it's clear that he never wanted a daughter and favors his Synthetic "son" David over her. It's sticking point that's made Meredith not only use a different last name, growing up in the shadow of David most of her life, but also one that makes her hope that the Prometheus mission is a failure - and that her father's precious Engineers either don't exist or turn out to not have the answers to immortality like he hopes. Weyland's desperate grasp for an extension to his life has caused him to fall and falter in her eyes. <br> Vickers is cold and obsessed with mitigating risk, causing some on board, who don't know she's Weyland's daughter, to think she might be an android herself. Meredith is there to follow her father's wishes, but also surprises him with her presence when he's awakened from cryosleep, as he only wanted David to accompany him on this mission, not her. Vickers passes the time leading up to New Year's Day 2094 by bossing people, having casual sex with Prometheus' pilot, Janek, and regretfully killing Dr. Charlie Holloway with a flamethrower when he's infected with the Engineers' Black Goo and begins mutating.<br> Towards the end, Vickers escapes the doomed Prometheus in an escape pod but once she touches back down on LV-223 she's crushed by the Engineer's ship as it falls from the sky. Janek (played by Idris Elba) Janek is the pilot of the USCSS Prometheus, and though he has a casual, flirty demeanor his first priority is the safety of his crew...and then eventually the safety of all life on Earth<br> Janek is never on any Away Mission to the Pyramid on LV-223, instead monitoring Pyramid dangers from inside Prometheus. When scientists Fifield and Milburn get lost and left behind in the Pyramid during a violent dust storm, Janek doesn't notice them being attacked by a Hammerpede, as he's away from his post and having sex with Weyland exec Meredith Vickers. Later, Janek heads down to the cargo bay to help his crew fight off and kill a mutated Fifield when he returns to the ship as a homicidal monster. <br> Confiding with Dr. Elizabeth Shaw, after the deaths of Fifield and Dr. Charlie Holloway, Janek informs her that he'll do whatever it takes to make sure nothing from LV-233 makes it back to Earth. When an Engineer is woken up and inadvertently set back on its mission to destroy Earth, Janek - with his co-pilots Chance and Ravel -- decide to use the full size of the Prometheus as a battering ram, launching their ship into the Engineer's ship, killing themselves and sending both ships crashing back down to LV-223.