Generation N: 0
012345678901234567890012345678901234567890
NAccbAaOccaaaaabccccbOAAaANObAbababcacccab-[0] = 4
NOONAONbAAcbbaabbbbccAOONaOacAOaccbaacacac-[1] = 4
AbabNcOcacaaaccbbbcccNcAbOaObcOabbbccbbcaa-[2] = 3
NNAcaNNANbccbcbbbacbcOacAabaNccbbbacbcbbcb-[3] = 6
ONbcaNNcAAccbcbbcaabcAaOaabbbAbaabccabbbca-[4] = 5
NObcNNaaNbabccccaccacNbcAbcNaAbacaaabcabab-[5] = 2
NOAacOcbbAacbaacaaabcAOacNaaabOcacccabbcbb-[6] = 4
ONaAcaNcOacbbbacbccacOcOAOcNcANcbcaabbbaba-[7] = 4
AOAONbAObbbbcbbacbbcbOONAbANAcbbcbabcccaab-[8] = 4
ANbbcbaaccaaaaccbabcbAOaaacObaObabccaacaba-[9] = 6
...
Generation N: 16
012345678901234567890012345678901234567890
AOcAcAOccNaaaaabccccbAbaabAbaOAababcacccab-[0] = 7
AaAcAaOAaOaaaaabccccbAOAbaObAObababcacccab-[1] = 7
OcOccAaOccaaaaabccccbAaAAbaabAaababcacccab-[2] = 7
AaOccAcAaOaaaaabccccbAOObAAOAObababcacccab-[3] = 7
OAaaaAaAaAaaaaabccccbObAObAAbaOababcacccab-[4] = 6
OAaOaAaAcAaaaaabccccbAAbObAObAAababcacccab-[5] = 7
AaAaAcAaOAaaaaabccccbObAObAAbaOababcacccab-[6] = 7
OccAcAaOccaaaaabccccbObAAOAObAAababcacccab-[7] = 6
OccAaOccAcaaaaabccccbObAAOAObAAababcacccab-[8] = 6
OccOccAaOcaaaaabccccbAaAAbaabAaababcacccab-[9] = 7
Generation N: 17
012345678901234567890012345678901234567890
OccOccAaOcaaaaabccccbAaAAbaabAaababcacccab-[0] = 7
AaOAaAaAcAaaaaabccccbObAObAAbaOababcacccab-[1] = 6
OAAaAaAcAaaaaaabccccbObAObAAbaOababcacccab-[2] = 6
AaOcOccAaOaaaaabccccbAaAAbaabAaababcacccab-[3] = 7
OaAaAOAaOaaaaaabccccbAAbObAObAAababcacccab-[4] = 7
OccAaOccOcaaaaabccccbAaAAbaabAaababcacccab-[5] = 7
AaAaAcAaOAaaaaabccccbAbObAObAAbababcacccab-[6] = 7
OcOccAaOccaaaaabccccbAAAaAAbaabababcacccab-[7] = 7
AcAaaOAaOaaaaaabccccbAAbObAObAAababcacccab-[8] = 8
AaOccAcAaOaaaaabccccbAAOAOObAAOababcacccab-[9] = 6
Figure 3.16. An initial population and its later descendants created via RIS transposition to solve the Majority (a, b, c) function problem. The chromosomes encode sub-ETs linked by OR. Note that none of the later descendants resemble their ancestors of generation 0. Note also the appearance of repetitive sequences in later generations. The perfect solution found by generation 17 (chromosome 8) and its mother (chromosome 5 of generation 16) are shown in blue. The event of transposition that led to this perfect solution can be seen in Figure 3.17.
It is worth emphasizing that this highly disruptive operator is, nevertheless, capable of forming simple, repetitive sequences like, for instance, the sequences (Aa)n or (Oc)n present in the later generations of
Figure 3.16. Interestingly, DNA is also full of repetitive sequences. In fact, in some eukaryotes more than 40% of DNA consists of small repetitive sequences. Most of these sequences are not even transcribed, but some genes are also interspersed with small islands of repetitive sequences.
Figure 3.17. Illustration of RIS transposition and its effects.
a) An event of RIS transposition with the transposon shown in bold. Note that a sequence with the length of the transposon is deleted at the end of the head of the target gene. Note also that the transposon became, in this case, only partially duplicated in the daughter chromosome; its other elements were deleted.
b) The sub-ETs encoded by the mother chromosome (before RIS transposition).
c) The sub-ETs encoded by the daughter chromosome (after RIS transposition). The transposon elements are shown in gray. Note that, in this case, root transposition changed drastically the
sub-ET1, shortening the mother sub-ET in eight nodes.