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Volgadraco

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Genus of ornithocheiroid pterosaur from the Late Cretaceous
Volgadraco
Temporal range: Late Cretaceous, Campanian
Speculative reconstruction of Volgadraco as an azhdarchid
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Clade: Pteranodontia
Genus: Volgadraco
Averianov et al., 2008[1]
Species:
V. bogolubovi
Binomial name
Volgadraco bogolubovi
Averianov et al., 2008[1]

Volgadraco ("Volga River dragon") is a genus of pteranodontian pterosaur from the Upper Cretaceous of European Russia.

Discovery and naming

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Volgadraco is known from lower beak (holotype SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like Azhdarcho and Bakonydraco and later Maastrichtian azhdarchids like Quetzalcoatlus . Volgadraco was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is V. bogolubovi, the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov.[1]

Classification

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Averianov and colleagues initially assigned Volgadraco to the family Azhdarchidae, and considered the earlier named genus Bogolubovia to be a nomen dubium that is potentially identical to Volgadraco.[1] In contrast, Longrich and colleagues in 2018 had recovered it within the family Nyctosauridae, the cladogram of which can be seen below:[2]

Pteranodontoidea
Pteranodontia
Pteranodontidae Nyctosauridae

Volgadraco bogolubovi

In 2020, Averianov and Arkhangelsky reconsidered both Bogolubovia and Volgadraco as members of the family Pteranodontidae.[3] In 2022, Averianov and Kurin considered both Volgadraco and Bogolubovia as valid pteranodontids based on a new specimen of Bogolubovia.[4] In the same year, however, Volgadraco was considered to be a pteranodontian both outside the Nyctosauridae and Pteranodontidae based on phylogenetic analysis, the cladogram of which can be seen below:[5]

Pteranodontia

Pteranodon sternbergi (Geosternbergia )

Volgadraco

Aponyctosauria
Nyctosauridae

Nyctosaurus grandis (Barbaridactylus )

Nyctosaurus nanus

Nyctosaurus gracilis

See also

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References

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  1. ^ a b c d Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (October 2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal. 42 (6): 634–642. Bibcode:2008PalJ...42..634A. doi:10.1134/S0031030108060099. S2CID 129558986.
  2. ^ Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3): e2001663. doi:10.1371/journal.pbio.2001663 . PMC 5849296 . PMID 29534059.
  3. ^ Alexander O. Averianov; Maxim S. Arkhangelsky (2020). "A large pteranodontid pterosaur from the Late Cretaceous of Eastern Europe". Geological Magazine. 158 (7): 1143–1155. doi:10.1017/S0016756820001119. S2CID 229441587.
  4. ^ Averianov AO, Kurin AS (2022). "A new specimen of pteranodontid pterosaur Bogolubovia orientalis from the Upper Cretaceous of Penza Province, Russia". Historical Biology: An International Journal of Paleobiology. 35 (8): 1288–1296. doi:10.1080/08912963.2022.2087522. S2CID 249728681.
  5. ^ Fernandes, Alexandra E.; Mateus, Octávio; Andres, Brian; Polcyn, Michael J.; Schulp, Anne S.; Gonçalves, António Olímpio; Jacobs, Louis L. (2022). "Pterosaurs from the Late Cretaceous of Angola". Diversity. 14 (9). 741. doi:10.3390/d14090741 . hdl:10362/145845 .
Avemetatarsalia
Pterosauria
    • see below↓
Preondactylia
Caviramidae?
Austriadraconidae
Raeticodactylidae
Eudimorphodontidae
Dimorphodontidae
Campylognathoididae
Rhamphorhynchidae
Scaphognathidae?
Pterodactylomorpha
    • see below↓
Darwinoptera
Wukongopteridae
Anurognathidae
Pterodactyloidea
Lophocratia
    • see below↓
Germanodactylidae
Gallodactylidae
Aurorazhdarchia
Aurorazhdarchidae
Ctenochasmatidae
Eupterodactyloidea
Ornithocheiroidea
    • see below↓
Dsungaripteridae
Thalassodromidae?
Tapejaridae
Dsungaripteromorpha?
Chaoyangopteridae
Azhdarchiformes
Alanqidae?
Azhdarchidae
Pteranodontoidea
    • see below↓
Pteranodontia
Pteranodontidae
Nyctosauromorpha
Aponyctosauria
Nyctosauridae
Lonchodectidae
Istiodactyliformes
Mimodactylidae
Istiodactylidae
Boreopteridae
Ornithocheiridae
Targaryendraconia?
Cimoliopteridae
Targaryendraconidae
Hamipteridae?
Anhangueridae

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