Bobcat data
Description
Example bobcat data for multimark package.
Format
The data are summarized in a 46x8 matrix containing observed encounter histories for 46 bobcats across 8 sampling occasions. Bobcats are bilaterially asymmetrical, and sampling was conducted using camera stations consisting of a single camera.
Because the left-side cannot be reconciled with the right-side, the two types of “marks” in this case are the pelage patterns on the left- and right-side of each individual. Encounter type 0 corresponds to non-detection, encounter type 1 corresponds to left-sided detection, encounter type 2 corresponds to right-sided detection.
Both-sided encounters were never observed in this dataset, hence the most appropriate multimark data type is data.type="never".
Source
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
See Also
Examples
data(bobcat)
Bobcat spatial capture-recapture data
Description
Example spatial bobcat data for multimark package.
Format
These spatial capture-recapture data with multiple mark types are summarized in a list of length 3 containing the following objects:
Enc.Mat is a 42 x (noccas*ntraps) matrix containing observed encounter histories for 42 bobcats across noccas=187 sampling occasions and ntraps=30 traps. The first 187 columns correspond to trap 1, the second 187 columns corresopond to trap 2, etc.
trapCoords is a matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate, and the second column the y-coordinate. The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating.
studyArea is a 3-column matrix containing the coordinates for the centroids of the contiguous grid of 1023 cells that define the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column indicates whether the cell is available habitat (=1) or not (=0). The grid cells are 0.65x0.65km resolution.
Bobcats are bilaterially asymmetrical, and sampling was conducted using camera stations consisting of a single camera. Because the left-side cannot be reconciled with the right-side, the two types of “marks” in this case are the pelage patterns on the left- and right-side of each individual. Encounter type 0 corresponds to non-detection, encounter type 1 corresponds to left-sided detection, encounter type 2 corresponds to right-sided detection.
Both-sided encounters were never observed in this dataset, hence the most appropriate multimark data type is data.type="never".
The first 15 rows of bobcatSCR$Enc.Mat correspond to individuals for which both the left and right sides were known because they were physically captured for telemetry deployments prior to sampling surveys. The encounter histories for these 15 individuals are therefore known with certainty and should be specified as such using the known argument in processdataSCR and/or multimarkClosedSCR (see example below).
These data were obtained from the R package SPIM (Augustine et al. 2017) and modified by projecting onto a regular rectangular grid consisting of square grid cells (as is required by the spatial capture-recapture models in multimark).
Details
We thank B. Augustine and co-authors for making these data publicly available in the SPIM package (Augustine et al. 2017).
Source
Augustine, B., Royle, J.A., Kelly, M., Satter, C., Alonso, R., Boydston, E. and Crooks, K. 2017. Spatial capture-recapture with partial identity: an application to camera traps. bioRxiv doi: https://doi.org/10.1101/056804
See Also
multimarkClosedSCR , processdataSCR
Examples
data(bobcatSCR)
#plot the traps and available habitat within the study area
plotSpatialData(trapCoords=bobcatSCR$trapCoords,studyArea=bobcatSCR$studyArea)
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
# Fit spatial model to tiger data
Enc.Mat <- bobcatSCR$Enc.Mat
trapCoords <- bobcatSCR$trapCoords
studyArea <- bobcatSCR$studyArea
# specify known encounter histories
known <- c(rep(1,15),rep(0,nrow(Enc.Mat)-15))
# specify prior bounds for sigma2_scr
sig_bounds <- c(0.1,max(diff(range(studyArea[,"x"])),diff(range(studyArea[,"y"]))))
mmsSCR <- processdataSCR(Enc.Mat,trapCoords,studyArea,known=known)
bobcatSCR.dot.type <- multimarkClosedSCR(mms=mmsSCR,iter=200,adapt=100,burnin=100,
sigma_bounds=sig_bounds)
summary(bobcatSCR.dot.type$mcmc)
Calculate population density estimates
Description
This function calculates posterior population density estimates from multimarkClosedSCR output as D = N/A, where D is density, N is abundance, and A is the area of available habitat within the study area.
Usage
getdensityClosedSCR(out)
Arguments
out
List of output returned by multimarkClosedSCR .
Value
An object of class mcmc.list containing the following:
D
Posterior samples for density.
Author(s)
Brett T. McClintock
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Run behavior model for simulated data with constant detection probability (i.e., mod.p=~c)
sim.data<-simdataClosedSCR()
Enc.Mat<-sim.data$Enc.Mat
trapCoords<-sim.data$spatialInputs$trapCoords
studyArea<-sim.data$spatialInputs$studyArea
example.dot <- multimarkClosedSCR(Enc.Mat,trapCoords,studyArea,mod.p=~1)
#Calculate capture and recapture probabilities
D <- getdensityClosedSCR(example.dot)
summary(D)
Calculate posterior capture and survival probabilities
Description
This function calculates posterior capture (p) and survival (\phi) probabilities for each sampling occasion from multimarkCJS output.
Usage
getprobsCJS(out, link = "probit")
Arguments
out
List of output returned by multimarkCJS
link
Link function for p and \phi. Must be "probit" or "logit". Note that multimarkCJS is currently implemented for the probit link only.
Value
An object of class mcmc.list containing the following:
p
Posterior samples for capture probability (p[c,t]) for each release cohort (c=1,\ldots,T-1) and sampling occasion (t=2,\ldots,T).
phi
Posterior samples for survival probability (\phi[c,k]) for each release cohort (c=1,\ldots,T-1) and interval (k=1,\ldots,T-1).
Author(s)
Brett T. McClintock
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Simulate open population data with temporal variation in survival
noccas <- 5
data <- simdataCJS(noccas=noccas, phibeta=rnorm(noccas-1,1.6,0.1))
#Fit open population model with temporal variation in survival
sim.time <- multimarkCJS(data$Enc.Mat,mod.phi=~time)
#Calculate capture and survival probabilities for each cohort and time
pphi <- getprobsCJS(sim.time)
summary(pphi)
Calculate posterior capture and recapture probabilities
Description
This function calculates posterior capture (p) and recapture (c) probabilities for each sampling occasion from multimarkClosed output.
Usage
getprobsClosed(out, link = "logit")
Arguments
out
List of output returned by multimarkClosed .
link
Link function for detection probability. Must be "logit" or "probit". Note that multimarkClosed is currently implemented for the logit link only.
Value
An object of class mcmc.list containing the following:
p
Posterior samples for capture probability (p) for each sampling occasion.
c
Posterior samples for recapture probability (c) for each sampling occasion.
Author(s)
Brett T. McClintock
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Run behavior model for bobcat data with constant detection probability (i.e., mod.p=~c)
bobcat.c <- multimarkClosed(bobcat,mod.p=~c)
#Calculate capture and recapture probabilities
pc <- getprobsClosed(bobcat.c)
summary(pc)
Calculate posterior capture and recapture probabilities
Description
This function calculates posterior spatial capture (p) and recapture (c) probabilities (at zero distance from an activity center) for each sampling occasion from multimarkClosedSCR output.
Usage
getprobsClosedSCR(out, link = "cloglog")
Arguments
out
List of output returned by multimarkClosedSCR .
link
Link function for detection probability. Must be "cloglog". Note that multimarkClosedSCR is currently implemented for the cloglog link only.
Value
An object of class mcmc.list containing the following:
p
Posterior samples for capture probability (p) for each sampling occasion (first index) and trap (second index).
c
Posterior samples for recapture probability (c) for each sampling occasion (first index) and trap (second index).
Author(s)
Brett T. McClintock
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Run behavior model for simulated data with constant detection probability (i.e., mod.p=~c)
sim.data<-simdataClosedSCR()
Enc.Mat<-sim.data$Enc.Mat
trapCoords<-sim.data$spatialInputs$trapCoords
studyArea<-sim.data$spatialInputs$studyArea
example.c <- multimarkClosedSCR(Enc.Mat,trapCoords,studyArea,mod.p=~c,
iter=1000,adapt=500,burnin=500)
#Calculate capture and recapture probabilities
pc <- getprobsClosedSCR(example.c)
summary(pc)
Fit open population survival models for “traditional” capture-mark-recapture data consisting of a single mark type
Description
This function fits Cormack-Jolly-Seber (CJS) open population models for survival probability (\phi) and capture probability (p) for “traditional” capture-mark-recapture data consisting of a single mark type. Using Bayesian analysis methods, Markov chain Monte Carlo (MCMC) is used to draw samples from the joint posterior distribution.
Usage
markCJS(
Enc.Mat,
covs = data.frame(),
mod.p = ~1,
mod.phi = ~1,
parms = c("pbeta", "phibeta"),
nchains = 1,
iter = 12000,
adapt = 1000,
bin = 50,
thin = 1,
burnin = 2000,
taccept = 0.44,
tuneadjust = 0.95,
proppbeta = 0.1,
propzp = 1,
propsigmap = 1,
propphibeta = 0.1,
propzphi = 1,
propsigmaphi = 1,
pbeta0 = 0,
pSigma0 = 1,
phibeta0 = 0,
phiSigma0 = 1,
l0p = 1,
d0p = 0.01,
l0phi = 1,
d0phi = 0.01,
initial.values = NULL,
link = "probit",
printlog = FALSE,
...
)
Arguments
Enc.Mat
A matrix of observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions. With a single mark type, encounter histories consist of only non-detections (0) and type 1 encounters (1).
covs
A data frame of temporal covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of sampling occasions. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
mod.p
Model formula for detection probability (p). For example, mod.p=~1 specifies no effects (i.e., intercept only), mod.p~time specifies temporal effects, mod.p~age specifies age effects, mod.p~h specifies individual heterogeneity, and mod.p~time+age specifies additive temporal and age effects.
mod.phi
Model formula for survival probability (\phi). For example, mod.phi=~1 specifies no effects (i.e., intercept only), mod.phi~time specifies temporal effects, mod.phi~age specifies age effects, mod.phi~h specifies individual heterogeneity, and mod.phi~time+age specifies additive temporal and age effects.
parms
A character vector giving the names of the parameters and latent variables to monitor. Possible parameters are probit-scale detection probability parameters ("pbeta" for p and "phibeta" for \phi), probit-scale individual heterogeneity variance terms ("sigma2_zp" for p and "sigma2_zphi" for \phi), and probit-scale individual effects ("zp" and "zphi"). Latent variable indicators for whether each individual was alive (1) or dead (0) during each sampling occasion ("q") and the log likelihood ("loglike") may also be monitored. Setting parms="all" monitors all possible parameters and latent variables.
nchains
The number of parallel MCMC chains for the model.
iter
The number of MCMC iterations.
adapt
Ignored; no adaptive phase is needed for "probit" link.
bin
Ignored; no adaptive phase is needed for "probit" link.
thin
Thinning interval for monitored parameters.
burnin
Number of burn-in iterations (0 <= burnin < iter).
taccept
Ignored; no adaptive phase is needed for "probit" link.
tuneadjust
Ignored; no adaptive phase is needed for "probit" link.
proppbeta
Ignored; no adaptive phase is needed for "probit" link.
propzp
Ignored; no adaptive phase is needed for "probit" link.
propsigmap
Ignored; no adaptive phase is needed for "probit" link.
propphibeta
Ignored; no adaptive phase is needed for "probit" link.
propzphi
Ignored; no adaptive phase is needed for "probit" link.
propsigmaphi
Ignored; no adaptive phase is needed for "probit" link.
pbeta0
Scaler or vector (of length k) specifying mean of pbeta ~ multivariateNormal(pbeta0, pSigma0) prior. If pbeta0 is a scaler, then this value is used for all j = 1, ..., k. Default is pbeta0 = 0.
pSigma0
Scaler or k x k matrix specifying covariance matrix of pbeta ~ multivariateNormal(pbeta0, pSigma0) prior. If pSigma0 is a scaler, then this value is used for all pSigma0[j,j] for j = 1, ..., k (with pSigma[j,l] = 0 for all j \ne l). Default is pSigma0 = 1.
phibeta0
Scaler or vector (of length k) specifying mean of phibeta ~ multivariateNormal(phibeta0, phiSigma0) prior. If phibeta0 is a scaler, then this value is used for all j = 1, ..., k. Default is phibeta0 = 0.
phiSigma0
Scaler or k x k matrix specifying covariance matrix of phibeta ~ multivariateNormal(phibeta0, phiSigma0) prior. If phiSigma0 is a scaler, then this value is used for all phiSigma0[j,j] for j = 1, ..., k (with phiSigma[j,l] = 0 for all j \ne l). Default is phiSigma0 = 1.
l0p
Specifies "shape" parameter for [sigma2_zp] ~ invGamma(l0p,d0p) prior. Default is l0p = 1.
d0p
Specifies "scale" parameter for [sigma2_zp] ~ invGamma(l0p,d0p) prior. Default is d0p = 0.01.
l0phi
Specifies "shape" parameter for [sigma2_zphi] ~ invGamma(l0phi,d0phi) prior. Default is l0phi = 1.
d0phi
Specifies "scale" parameter for [sigma2_zphi] ~ invGamma(l0phi,d0phi) prior. Default is d0phi = 0.01.
initial.values
OOptional list of nchain list(s) specifying initial values for "pbeta", "phibeta", "sigma2_zp", "sigma2_zphi", "zp", "zphi", and "q". Default is initial.values = NULL, which causes initial values to be generated automatically.
link
Link function for survival and capture probabilities. Only probit link is currently implemented.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
...
Additional "parameters" arguments for specifying mod.p and mod.phi. See RMark::make.design.data .
Details
The first time markCJS (or markClosed ) is called, it will likely produce a firewall warning alerting users that R has requested the ability to accept incoming network connections. Incoming network connections are required to use parallel processing as implemented in multimarkCJS. Note that setting parms="all" is required for any markCJS model output to be used in multimodelCJS .
Value
A list containing the following:
mcmc
Markov chain Monte Carlo object of class mcmc.list .
mod.p
Model formula for detection probability (as specified by mod.p above).
mod.phi
Model formula for survival probability (as specified by mod.phi above).
mod.delta
Formula always NULL; only for internal use in multimodelCJS .
DM
A list of design matrices for detection and survival probability respectively generated by mod.p and mod.phi, where DM$p is the design matrix for capture probability (p) and DM$phi is the design matrix for survival probability (\phi).
initial.values
A list containing the parameter and latent variable values at iteration iter for each chain. Values are provided for "pbeta", "phibeta", "sigma2_zp", "sigma2_zphi", "zp", "zphi", and "q".
mms
An object of class multimarksetup
Author(s)
Brett T. McClintock
See Also
Examples
# These examples are excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Simulate open population data using defaults
data <- simdataCJS(delta_1=1,delta_2=0)$Enc.Mat
#Fit default open population model
sim.dot <- markCJS(data)
#Posterior summary for monitored parameters
summary(sim.dot$mcmc)
plot(sim.dot$mcmc)
#Fit ``age'' model with 2 age classes (e.g., juvenile and adult) for survival
#using 'parameters' and 'right' arguments from RMark::make.design.data
sim.age <- markCJS(data,mod.phi=~age,
parameters=list(Phi=list(age.bins=c(0,1,4))),right=FALSE)
summary(getprobsCJS(sim.age))
Fit closed population abundance models for “traditional” capture-mark-recapture data consisting of a single mark type
Description
This function fits closed population abundance models for “traditional” capture-mark-recapture data consisting of a single mark type using Bayesian analysis methods. Markov chain Monte Carlo (MCMC) is used to draw samples from the joint posterior distribution.
Usage
markClosed(
Enc.Mat,
covs = data.frame(),
mod.p = ~1,
parms = c("pbeta", "N"),
nchains = 1,
iter = 12000,
adapt = 1000,
bin = 50,
thin = 1,
burnin = 2000,
taccept = 0.44,
tuneadjust = 0.95,
proppbeta = 0.1,
propzp = 1,
propsigmap = 1,
npoints = 500,
a = 25,
mu0 = 0,
sigma2_mu0 = 1.75,
initial.values = NULL,
printlog = FALSE,
...
)
Arguments
Enc.Mat
A matrix of observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions. With a single mark type, encounter histories consist of only non-detections (0) and type 1 encounters (1).
covs
A data frame of temporal covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of sampling occasions. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
mod.p
Model formula for detection probability. For example, mod.p=~1 specifies no effects (i.e., intercept only), mod.p~time specifies temporal effects, mod.p~c specifies behavioral reponse (i.e., trap "happy" or "shy"), mod.p~h specifies individual heterogeneity, and mod.p~time+c specifies additive temporal and behavioral effects.
parms
A character vector giving the names of the parameters and latent variables to monitor. Possible parameters are logit-scale detection probability parameters ("pbeta"), population abundance ("N"), logit-scale individual heterogeneity variance term ("sigma2_zp"), and logit-scale individual effects ("zp"). The log posterior density ("logPosterior") may also be monitored. Setting parms="all" monitors all possible parameters and latent variables.
nchains
The number of parallel MCMC chains for the model.
iter
The number of MCMC iterations.
adapt
The number of iterations for proposal distribution adaptation. If adapt = 0 then no adaptation occurs.
bin
Bin length for calculating acceptance rates during adaptive phase (0 < bin <= iter).
thin
Thinning interval for monitored parameters.
burnin
Number of burn-in iterations (0 <= burnin < iter).
taccept
Target acceptance rate during adaptive phase (0 < taccept <= 1). Acceptance rate is monitored every bin iterations. Default is taccept = 0.44.
tuneadjust
Adjustment term during adaptive phase (0 < tuneadjust <= 1). If acceptance rate is less than taccept, then proposal term (proppbeta, propzp, or propsigmap) is multiplied by tuneadjust. If acceptance rate is greater than or equal to taccept, then proposal term is divided by tuneadjust. Default is tuneadjust = 0.95.
proppbeta
Scaler or vector (of length k) specifying the initial standard deviation of the Normal(pbeta[j], proppbeta[j]) proposal distribution. If proppbeta is a scaler, then this value is used for all j = 1, ..., k. Default is proppbeta = 0.1.
propzp
Scaler or vector (of length M) specifying the initial standard deviation of the Normal(zp[i], propzp[i]) proposal distribution. If propzp is a scaler, then this value is used for all i = 1, ..., M individuals. Default is propzp = 1.
propsigmap
Scaler specifying the initial Gamma(shape = 1/propsigmap, scale = sigma_zp * propsigmap) proposal distribution for sigma_zp = sqrt(sigma2_zp). Default is propsigmap=1.
npoints
Number of Gauss-Hermite quadrature points to use for numerical integration. Accuracy increases with number of points, but so does computation time.
a
Scale parameter for [sigma_z] ~ half-Cauchy(a) prior for the individual hetegeneity term sigma_zp = sqrt(sigma2_zp). Default is “uninformative” a = 25.
mu0
Scaler or vector (of length k) specifying mean of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is mu0 = 0.
sigma2_mu0
Scaler or vector (of length k) specifying variance of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If sigma2_mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is sigma2_mu0 = 1.75.
initial.values
Optional list of nchain list(s) specifying initial values for "pbeta", "zp", "sigma2_zp", and "N". Default is initial.values = NULL, which causes initial values to be generated automatically.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
...
Additional "parameters" arguments for specifying mod.p. See make.design.data .
Details
The first time markClosed (or markCJS ) is called, it will likely produce a firewall warning alerting users that R has requested the ability to accept incoming network connections. Incoming network connections are required to use parallel processing as implemented in markClosed. Note that setting parms="all" is required for any markClosed model output to be used in multimodelClosed .
Value
A list containing the following:
mcmc
Markov chain Monte Carlo object of class mcmc.list .
mod.p
Model formula for detection probability (as specified by mod.p above).
mod.delta
Formula always NULL; only for internal use in multimodelClosed .
DM
A list of design matrices for detection probability generated for model mod.p, where DM$p is the design matrix for initial capture probability (p) and DM$c is the design matrix for recapture probability (c).
initial.values
A list containing the parameter and latent variable values at iteration iter for each chain. Values are provided for "pbeta", "zp", "sigma2_zp", and "N".
mms
An object of class multimarksetup
Author(s)
Brett T. McClintock
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Run single chain using the default model for simulated ``traditional'' data
data<-simdataClosed(delta_1=1,delta_2=0)$Enc.Mat
sim.dot<-markClosed(data)
#Posterior summary for monitored parameters
summary(sim.dot$mcmc)
plot(sim.dot$mcmc)
Fit spatial population abundance models for “traditional” capture-mark-recapture data consisting of a single mark type
Description
This function fits spatial population abundance models for “traditional” capture-mark-recapture data consisting of a single mark type using Bayesian analysis methods. Markov chain Monte Carlo (MCMC) is used to draw samples from the joint posterior distribution.
Usage
markClosedSCR(
Enc.Mat,
trapCoords,
studyArea = NULL,
buffer = NULL,
ncells = 1024,
covs = data.frame(),
mod.p = ~1,
detection = "half-normal",
parms = c("pbeta", "N"),
nchains = 1,
iter = 12000,
adapt = 1000,
bin = 50,
thin = 1,
burnin = 2000,
taccept = 0.44,
tuneadjust = 0.95,
proppbeta = 0.1,
propsigma = 1,
propcenter = NULL,
sigma_bounds = NULL,
mu0 = 0,
sigma2_mu0 = 1.75,
initial.values = NULL,
scalemax = 10,
printlog = FALSE,
...
)
Arguments
Enc.Mat
A matrix containing the observed encounter histories with rows corresponding to individuals and (ntraps*noccas) columns corresponding to traps and sampling occasions. The first noccas columns correspond to trap 1, the second noccas columns corresopond to trap 2, etc.
trapCoords
A matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate (“x”), and the second column the y-coordinate (“y”). The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating. Ignored unless mms=NULL.
studyArea
is a 3-column matrix containing the coordinates for the centroids a contiguous grid of cells that define the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns (“x” and “y”) indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column (“avail”) indicates whether the cell is available habitat (=1) or not (=0). All cells must have the same resolution. If studyArea=NULL (the default) and mms=NULL, then a square study area grid composed of ncells cells of available habitat is drawn around the bounding box of trapCoords based on buffer. Ignored unless mms=NULL.
Note that rows should be ordered by raster cell order (raster cell numbers start at 1 in the upper left corner, and increase from left to right, and then from top to bottom).
buffer
A scaler in same units as trapCoords indicating the buffer around the bounding box of trapCoords for defining the study area when studyArea=NULL. Ignored unless studyArea=NULL.
ncells
The number of grid cells in the study area when studyArea=NULL. The square root of ncells must be a whole number. Default is ncells=1024. Ignored unless studyArea=NULL and mms=NULL.
covs
A data frame of time- and/or trap-dependent covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of traps times the number of sampling occasions (ntraps*noccas), where the first noccas rows correspond to trap 1, the noccas rows correspond to trap 2, etc. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
mod.p
Model formula for detection probability. For example, mod.p=~1 specifies no effects (i.e., intercept only), mod.p~time specifies temporal effects, mod.p~c specifies behavioral reponse (i.e., trap "happy" or "shy"), mod.p~trap specifies trap effects, and mod.p~time+c specifies additive temporal and behavioral effects.
detection
Model for detection probability as a function of distance from activity centers . Must be "half-normal" (of the form \exp{(-d^2 / (2*\sigma^2))}, where d is distance) or "exponential" (of the form \exp{(-d / \lambda)}).
parms
A character vector giving the names of the parameters and latent variables to monitor. Possible parameters are cloglog-scale detection probability parameters ("pbeta"), population abundance ("N"), and cloglog-scale distance term for the detection function ("sigma2_scr" when detection=``half-normal'' or "lambda" when detection=``exponential''). Individual activity centers ("centers") and the log posterior density ("logPosterior") may also be monitored. Setting parms="all" monitors all possible parameters and latent variables.
nchains
The number of parallel MCMC chains for the model.
iter
The number of MCMC iterations.
adapt
The number of iterations for proposal distribution adaptation. If adapt = 0 then no adaptation occurs.
bin
Bin length for calculating acceptance rates during adaptive phase (0 < bin <= iter).
thin
Thinning interval for monitored parameters.
burnin
Number of burn-in iterations (0 <= burnin < iter).
taccept
Target acceptance rate during adaptive phase (0 < taccept <= 1). Acceptance rate is monitored every bin iterations. Default is taccept = 0.44.
tuneadjust
Adjustment term during adaptive phase (0 < tuneadjust <= 1). If acceptance rate is less than taccept, then proposal term (proppbeta or propsigma) is multiplied by tuneadjust. If acceptance rate is greater than or equal to taccept, then proposal term is divided by tuneadjust. Default is tuneadjust = 0.95.
proppbeta
Scaler or vector (of length k) specifying the initial standard deviation of the Normal(pbeta[j], proppbeta[j]) proposal distribution. If proppbeta is a scaler, then this value is used for all j = 1, ..., k. Default is proppbeta = 0.1.
propsigma
Scaler specifying the initial Gamma(shape = 1/propsigma, scale = sigma_scr * propsigma) proposal distribution for sigma_scr = sqrt(sigma2_scr). Default is propsigma=1.
propcenter
Scaler specifying the neighborhood distance when proposing updates to activity centers. When propcenter=NULL (the default), then propcenter = a*10, where a is the cell size for the study area grid, and each cell has (at most) approximately 300 neighbors.
sigma_bounds
Positive vector of length 2 for the lower and upper bounds for the [sigma_scr] ~ Uniform(sigma_bounds[1], sigma_bounds[2]) (or [sqrt(lambda)] when detection=``exponential'') prior for the detection function term sigma_scr = sqrt(sigma2_scr) (or sqrt(lambda)). When sigma_bounds = NULL (the default), then sigma_bounds = c(1.e-6,max(diff(range(studyArea[,"x"])),diff(range(studyArea[,"y"])))).
mu0
Scaler or vector (of length k) specifying mean of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is mu0 = 0.
sigma2_mu0
Scaler or vector (of length k) specifying variance of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If sigma2_mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is sigma2_mu0 = 1.75.
initial.values
Optional list of nchain list(s) specifying initial values for "pbeta", "N", "sigma2_scr", and "centers". Default is initial.values = NULL, which causes initial values to be generated automatically.
scalemax
Upper bound for internal re-scaling of grid cell centroid coordinates. Default is scalemax=10, which re-scales the centroids to be between 0 and 10. Re-scaling is done internally to avoid numerical overflows during model fitting.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
...
Additional "parameters" arguments for specifying mod.p. See make.design.data .
Details
The first time markClosedSCR is called, it will likely produce a firewall warning alerting users that R has requested the ability to accept incoming network connections. Incoming network connections are required to use parallel processing as implemented in markClosed. Note that setting parms="all" is required for any markClosed model output to be used in multimodelClosed .
Value
A list containing the following:
mcmc
Markov chain Monte Carlo object of class mcmc.list .
mod.p
Model formula for detection probability (as specified by mod.p above).
mod.delta
Formula always NULL; only for internal use in multimodelClosedSCR .
mod.det
Model formula for detection function (as specified by detection above).
DM
A list of design matrices for detection probability generated for model mod.p, where DM$p is the design matrix for initial capture probability (p) and DM$c is the design matrix for recapture probability (c).
initial.values
A list containing the parameter and latent variable values at iteration iter for each chain. Values are provided for "pbeta", "N", "sigma2_scr", and "centers".
mms
An object of class multimarkSCRsetup
Author(s)
Brett T. McClintock
References
Gopalaswamy, A.M., Royle, J.A., Hines, J.E., Singh, P., Jathanna, D., Kumar, N. and Karanth, K.U. 2012. Program SPACECAP: software for estimating animal density using spatially explicit capture-recapture models. Methods in Ecology and Evolution 3:1067-1072.
King, R., McClintock, B. T., Kidney, D., and Borchers, D. L. 2016. Capture-recapture abundance estimation using a semi-complete data likelihood approach. The Annals of Applied Statistics 10: 264-285
Royle, J.A., Karanth, K.U., Gopalaswamy, A.M. and Kumar, N.S. 2009. Bayesian inference in camera trapping studies for a class of spatial capture-recapture models. Ecology 90: 3233-3244.
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Run single chain using the default model for ``traditional'' tiger data of Royle et al (2009)
Enc.Mat<-tiger$Enc.Mat
trapCoords<-tiger$trapCoords
studyArea<-tiger$studyArea
tiger.dot<-markClosedSCR(Enc.Mat,trapCoords,studyArea,iter=100,adapt=50,burnin=50)
#Posterior summary for monitored parameters
summary(tiger.dot$mcmc)
plot(tiger.dot$mcmc)
Fit open population survival models for capture-mark-recapture data consisting of multiple non-invasive marks
Description
This function fits Cormack-Jolly-Seber (CJS) open population models for survival probability (\phi) and capture probability (p) from capture-mark-recapture data consisting of multiple non-invasive marks. Using Bayesian analysis methods, Markov chain Monte Carlo (MCMC) is used to draw samples from the joint posterior distribution.
Usage
multimarkCJS(
Enc.Mat,
data.type = "never",
covs = data.frame(),
mms = NULL,
mod.p = ~1,
mod.phi = ~1,
mod.delta = ~type,
parms = c("pbeta", "phibeta", "delta"),
nchains = 1,
iter = 12000,
adapt = 1000,
bin = 50,
thin = 1,
burnin = 2000,
taccept = 0.44,
tuneadjust = 0.95,
proppbeta = 0.1,
propzp = 1,
propsigmap = 1,
propphibeta = 0.1,
propzphi = 1,
propsigmaphi = 1,
maxnumbasis = 1,
pbeta0 = 0,
pSigma0 = 1,
phibeta0 = 0,
phiSigma0 = 1,
l0p = 1,
d0p = 0.01,
l0phi = 1,
d0phi = 0.01,
a0delta = 1,
a0alpha = 1,
b0alpha = 1,
a0psi = 1,
b0psi = 1,
initial.values = NULL,
known = integer(),
link = "probit",
printlog = FALSE,
...
)
Arguments
Enc.Mat
A matrix of observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions (ignored unless mms=NULL).
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
covs
A data frame of temporal covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of sampling occasions. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
mms
An optional object of class multimarksetup-class; if NULL it is created. See processdata .
mod.p
Model formula for detection probability (p). For example, mod.p=~1 specifies no effects (i.e., intercept only), mod.p~time specifies temporal effects, mod.p~age specifies age effects, mod.p~h specifies individual heterogeneity, and mod.p~time+age specifies additive temporal and age effects.
mod.phi
Model formula for survival probability (\phi). For example, mod.phi=~1 specifies no effects (i.e., intercept only), mod.phi~time specifies temporal effects, mod.phi~age specifies age effects, mod.phi~h specifies individual heterogeneity, and mod.phi~time+age specifies additive temporal and age effects.
mod.delta
Model formula for conditional probabilities of type 1 (delta_1) and type 2 (delta_2) encounters, given detection. Currently only mod.delta=~1 (i.e., \delta_1 = \delta_2) and mod.delta=~type (i.e., \delta_1 \ne \delta_2) are implemented.
parms
A character vector giving the names of the parameters and latent variables to monitor. Possible parameters are probit-scale detection probability parameters ("pbeta" for p and "phibeta" for \phi), conditional probability of type 1 or type 2 encounter, given detection ("delta)", probability of simultaneous type 1 and type 2 detection, given both types encountered ("alpha"), probit-scale individual heterogeneity variance terms ("sigma2_zp" for p and "sigma2_zphi" for \phi), probit-scale individual effects ("zp" and "zphi"), and the probability that a randomly selected individual from the M = nrow(Enc.Mat) observed individuals belongs to the n unique individuals encountered at least once ("psi"). Individual encounter history indices ("H"), latent variable indicators for whether each individual was alive (1) or dead (0) during each sampling occasion ("q"), and the log likelihood ("loglike") may also be monitored. Setting parms="all" monitors all possible parameters and latent variables.
nchains
The number of parallel MCMC chains for the model.
iter
The number of MCMC iterations.
adapt
Ignored; no adaptive phase is needed for "probit" link.
bin
Ignored; no adaptive phase is needed for "probit" link.
thin
Thinning interval for monitored parameters.
burnin
Number of burn-in iterations (0 <= burnin < iter).
taccept
Ignored; no adaptive phase is needed for "probit" link.
tuneadjust
Ignored; no adaptive phase is needed for "probit" link.
proppbeta
Ignored; no adaptive phase is needed for "probit" link.
propzp
Ignored; no adaptive phase is needed for "probit" link.
propsigmap
Ignored; no adaptive phase is needed for "probit" link.
propphibeta
Ignored; no adaptive phase is needed for "probit" link.
propzphi
Ignored; no adaptive phase is needed for "probit" link.
propsigmaphi
Ignored; no adaptive phase is needed for "probit" link.
maxnumbasis
Maximum number of basis vectors to use when proposing latent history frequency updates. Default is maxnumbasis = 1, but higher values can potentially improve mixing.
pbeta0
Scaler or vector (of length k) specifying mean of pbeta ~ multivariateNormal(pbeta0, pSigma0) prior. If pbeta0 is a scaler, then this value is used for all j = 1, ..., k. Default is pbeta0 = 0.
pSigma0
Scaler or k x k matrix specifying covariance matrix of pbeta ~ multivariateNormal(pbeta0, pSigma0) prior. If pSigma0 is a scaler, then this value is used for all pSigma0[j,j] for j = 1, ..., k (with pSigma[j,l] = 0 for all j \ne l). Default is pSigma0 = 1.
phibeta0
Scaler or vector (of length k) specifying mean of phibeta ~ multivariateNormal(phibeta0, phiSigma0) prior. If phibeta0 is a scaler, then this value is used for all j = 1, ..., k. Default is phibeta0 = 0.
phiSigma0
Scaler or k x k matrix specifying covariance matrix of phibeta ~ multivariateNormal(phibeta0, phiSigma0) prior. If phiSigma0 is a scaler, then this value is used for all phiSigma0[j,j] for j = 1, ..., k (with phiSigma[j,l] = 0 for all j \ne l). Default is phiSigma0 = 1.
l0p
Specifies "shape" parameter for [sigma2_zp] ~ invGamma(l0p,d0p) prior. Default is l0p = 1.
d0p
Specifies "scale" parameter for [sigma2_zp] ~ invGamma(l0p,d0p) prior. Default is d0p = 0.01.
l0phi
Specifies "shape" parameter for [sigma2_zphi] ~ invGamma(l0phi,d0phi) prior. Default is l0phi = 1.
d0phi
Specifies "scale" parameter for [sigma2_zphi] ~ invGamma(l0phi,d0phi) prior. Default is d0phi = 0.01.
a0delta
Scaler or vector (of length d) specifying the prior for the conditional (on detection) probability of type 1 (delta_1), type 2 (delta_2), and both type 1 and type 2 encounters (1-delta_1-delta_2). If a0delta is a scaler, then this value is used for all a0delta[j] for j = 1, ..., d. For mod.delta=~type, d=3 with [delta_1, delta_2, 1-delta_1-delta_2] ~ Dirichlet(a0delta) prior. For mod.delta=~1, d=2 with [tau] ~ Beta(a0delta[1],a0delta[2]) prior, where (delta_1,delta_2,1-delta_1-delta_2) = (tau/2,tau/2,1-tau). See McClintock et al. (2013) for more details.
a0alpha
Specifies "shape1" parameter for [alpha] ~ Beta(a0alpha, b0alpha) prior. Only applicable when data.type = "sometimes". Default is a0alpha = 1. Note that when a0alpha = 1 and b0alpha = 1, then [alpha] ~ Unif(0,1).
b0alpha
Specifies "shape2" parameter for [alpha] ~ Beta(a0alpha, b0alpha) prior. Only applicable when data.type = "sometimes". Default is b0alpha = 1. Note that when a0alpha = 1 and b0alpha = 1, then [alpha] ~ Unif(0,1).
a0psi
Specifies "shape1" parameter for [psi] ~ Beta(a0psi,b0psi) prior. Default is a0psi = 1.
b0psi
Specifies "shape2" parameter for [psi] ~ Beta(a0psi,b0psi) prior. Default is b0psi = 1.
initial.values
Optional list of nchain list(s) specifying initial values for parameters and latent variables. Default is initial.values = NULL, which causes initial values to be generated automatically. In addition to the parameters ("pbeta", "phibeta", "delta_1", "delta_2", "alpha", "sigma2_zp", "sigma2_zphi", "zp", "zphi", and "psi"), initial values can be specified for the initial latent history frequencies ("x"), initial individual encounter history indices ("H"), and initial latent variable indicators for whether each individual was alive (1) or dead (0) during each sampling occasion ("q").
known
Optional integer vector indicating whether the encounter history of an individual is known with certainty (i.e., the observed encounter history is the true encounter history). Encounter histories with at least one type 4 encounter are automatically assumed to be known, and known does not need to be specified unless there exist encounter histories that do not contain a type 4 encounter that happen to be known with certainty (e.g., from independent telemetry studies). If specified, known = c(v_1,v_2,...,v_M) must be a vector of length M = nrow(Enc.Mat) where v_i = 1 if the encounter history for individual i is known (v_i = 0 otherwise). Note that known all-zero encounter histories (e.g., ‘000’) are ignored.
link
Link function for survival and capture probabilities. Only probit link is currently implemented.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
...
Additional "parameters" arguments for specifying mod.p and mod.phi. See RMark::make.design.data .
Details
The first time multimarkCJS (or multimarkClosed ) is called, it will likely produce a firewall warning alerting users that R has requested the ability to accept incoming network connections. Incoming network connections are required to use parallel processing as implemented in multimarkCJS. Note that setting parms="all" is required for any multimarkCJS model output to be used in multimodelCJS .
Value
A list containing the following:
mcmc
Markov chain Monte Carlo object of class mcmc.list .
mod.p
Model formula for detection probability (as specified by mod.p above).
mod.phi
Model formula for survival probability (as specified by mod.phi above).
mod.delta
Formula always NULL; only for internal use in multimodelCJS .
DM
A list of design matrices for detection and survival probability respectively generated by mod.p and mod.phi, where DM$p is the design matrix for capture probability (p) and DM$phi is the design matrix for survival probability (\phi).
initial.values
A list containing the parameter and latent variable values at iteration iter for each chain. Values are provided for "pbeta", "phibeta", "delta_1", "delta_2", "alpha", "sigma2_zp" "sigma2_zphi", "zp", "zphi", "psi", "x", "H", and "q".
mms
An object of class multimarksetup
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
McClintock, B. T., Bailey, L. L., Dreher, B. P., and Link, W. A. 2014. Probit models for capture-recapture data subject to imperfect detection, individual heterogeneity and misidentification. The Annals of Applied Statistics 8: 2461-2484.
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Simulate open population data using defaults
data <- simdataCJS()
#Fit default open population model
sim.dot <- multimarkCJS(data$Enc.Mat)
#Posterior summary for monitored parameters
summary(sim.dot$mcmc)
plot(sim.dot$mcmc)
#' #Fit ``age'' model with 2 age classes (e.g., juvenile and adult) for survival
#using 'parameters' and 'right' arguments from RMark::make.design.data
sim.age <- multimarkCJS(data$Enc.Mat,mod.phi=~age,
parameters=list(Phi=list(age.bins=c(0,1,4))),right=FALSE)
summary(getprobsCJS(sim.age))
Fit closed population abundance models for capture-mark-recapture data consisting of multiple non-invasive marks
Description
This function fits closed population abundance models for capture-mark-recapture data consisting of multiple non-invasive marks using Bayesian analysis methods. Markov chain Monte Carlo (MCMC) is used to draw samples from the joint posterior distribution.
Usage
multimarkClosed(
Enc.Mat,
data.type = "never",
covs = data.frame(),
mms = NULL,
mod.p = ~1,
mod.delta = ~type,
parms = c("pbeta", "delta", "N"),
nchains = 1,
iter = 12000,
adapt = 1000,
bin = 50,
thin = 1,
burnin = 2000,
taccept = 0.44,
tuneadjust = 0.95,
proppbeta = 0.1,
propzp = 1,
propsigmap = 1,
npoints = 500,
maxnumbasis = 1,
a0delta = 1,
a0alpha = 1,
b0alpha = 1,
a = 25,
mu0 = 0,
sigma2_mu0 = 1.75,
a0psi = 1,
b0psi = 1,
initial.values = NULL,
known = integer(),
printlog = FALSE,
...
)
Arguments
Enc.Mat
A matrix of observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions (ignored unless mms=NULL).
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
covs
A data frame of temporal covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of sampling occasions. Covariate names cannot be "time", "c", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
mms
An optional object of class multimarksetup-class; if NULL it is created. See processdata .
mod.p
Model formula for detection probability. For example, mod.p=~1 specifies no effects (i.e., intercept only), mod.p~time specifies temporal effects, mod.p~c specifies behavioral reponse (i.e., trap "happy" or "shy"), mod.p~h specifies individual heterogeneity, and mod.p~time+c specifies additive temporal and behavioral effects.
mod.delta
Model formula for conditional probabilities of type 1 (delta_1) and type 2 (delta_2) encounters, given detection. Currently only mod.delta=~1 (i.e., \delta_1 = \delta_2) and mod.delta=~type (i.e., \delta_1 \ne \delta_2) are implemented.
parms
A character vector giving the names of the parameters and latent variables to monitor. Possible parameters are logit-scale detection probability parameters ("pbeta"), population abundance ("N"), conditional probability of type 1 or type 2 encounter, given detection ("delta)", probability of simultaneous type 1 and type 2 detection, given both types encountered ("alpha"), logit-scale individual heterogeneity variance term ("sigma2_zp"), logit-scale individual effects ("zp"), and the probability that a randomly selected individual from the M = nrow(Enc.Mat) observed individuals belongs to the n unique individuals encountered at least once ("psi"). Individual encounter history indices ("H") and the log posterior density ("logPosterior") may also be monitored. Setting parms="all" monitors all possible parameters and latent variables.
nchains
The number of parallel MCMC chains for the model.
iter
The number of MCMC iterations.
adapt
The number of iterations for proposal distribution adaptation. If adapt = 0 then no adaptation occurs.
bin
Bin length for calculating acceptance rates during adaptive phase (0 < bin <= iter).
thin
Thinning interval for monitored parameters.
burnin
Number of burn-in iterations (0 <= burnin < iter).
taccept
Target acceptance rate during adaptive phase (0 < taccept <= 1). Acceptance rate is monitored every bin iterations. Default is taccept = 0.44.
tuneadjust
Adjustment term during adaptive phase (0 < tuneadjust <= 1). If acceptance rate is less than taccept, then proposal term (proppbeta, propzp, or propsigmap) is multiplied by tuneadjust. If acceptance rate is greater than or equal to taccept, then proposal term is divided by tuneadjust. Default is tuneadjust = 0.95.
proppbeta
Scaler or vector (of length k) specifying the initial standard deviation of the Normal(pbeta[j], proppbeta[j]) proposal distribution. If proppbeta is a scaler, then this value is used for all j = 1, ..., k. Default is proppbeta = 0.1.
propzp
Scaler or vector (of length M) specifying the initial standard deviation of the Normal(zp[i], propzp[i]) proposal distribution. If propzp is a scaler, then this value is used for all i = 1, ..., M individuals. Default is propzp = 1.
propsigmap
Scaler specifying the initial Gamma(shape = 1/propsigmap, scale = sigma_zp * propsigmap) proposal distribution for sigma_zp = sqrt(sigma2_zp). Default is propsigmap=1.
npoints
Number of Gauss-Hermite quadrature points to use for numerical integration. Accuracy increases with number of points, but so does computation time.
maxnumbasis
Maximum number of basis vectors to use when proposing latent history frequency updates. Default is maxnumbasis = 1, but higher values can potentially improve mixing.
a0delta
Scaler or vector (of length d) specifying the prior for the conditional (on detection) probability of type 1 (delta_1), type 2 (delta_2), and both type 1 and type 2 encounters (1-delta_1-delta_2). If a0delta is a scaler, then this value is used for all a0delta[j] for j = 1, ..., d. For mod.delta=~type, d=3 with [delta_1, delta_2, 1-delta_1-delta_2] ~ Dirichlet(a0delta) prior. For mod.delta=~1, d=2 with [tau] ~ Beta(a0delta[1],a0delta[2]) prior, where (delta_1,delta_2,1-delta_1-delta_2) = (tau/2,tau/2,1-tau). See McClintock et al. (2013) for more details.
a0alpha
Specifies "shape1" parameter for [alpha] ~ Beta(a0alpha, b0alpha) prior. Only applicable when data.type = "sometimes". Default is a0alpha = 1. Note that when a0alpha = 1 and b0alpha = 1, then [alpha] ~ Unif(0,1).
b0alpha
Specifies "shape2" parameter for [alpha] ~ Beta(a0alpha, b0alpha) prior. Only applicable when data.type = "sometimes". Default is b0alpha = 1. Note that when a0alpha = 1 and b0alpha = 1, then [alpha] ~ Unif(0,1).
a
Scale parameter for [sigma_z] ~ half-Cauchy(a) prior for the individual hetegeneity term sigma_zp = sqrt(sigma2_zp). Default is “uninformative” a = 25.
mu0
Scaler or vector (of length k) specifying mean of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is mu0 = 0.
sigma2_mu0
Scaler or vector (of length k) specifying variance of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If sigma2_mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is sigma2_mu0 = 1.75.
a0psi
Specifies "shape1" parameter for [psi] ~ Beta(a0psi,b0psi) prior. Default is a0psi = 1.
b0psi
Specifies "shape2" parameter for [psi] ~ Beta(a0psi,b0psi) prior. Default is b0psi = 1.
initial.values
Optional list of nchain list(s) specifying initial values for parameters and latent variables. Default is initial.values = NULL, which causes initial values to be generated automatically. In addition to the parameters ("pbeta", "N", "delta_1", "delta_2", "alpha", "sigma2_zp", "zp", and "psi"), initial values can be specified for the initial latent history frequencies ("x") and initial individual encounter history indices ("H").
known
Optional integer vector indicating whether the encounter history of an individual is known with certainty (i.e., the observed encounter history is the true encounter history). Encounter histories with at least one type 4 encounter are automatically assumed to be known, and known does not need to be specified unless there exist encounter histories that do not contain a type 4 encounter that happen to be known with certainty (e.g., from independent telemetry studies). If specified, known = c(v_1,v_2,...,v_M) must be a vector of length M = nrow(Enc.Mat) where v_i = 1 if the encounter history for individual i is known (v_i = 0 otherwise). Note that known all-zero encounter histories (e.g., ‘000’) are ignored.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
...
Additional "parameters" arguments for specifying mod.p. See make.design.data .
Details
The first time multimarkClosed (or multimarkCJS ) is called, it will likely produce a firewall warning alerting users that R has requested the ability to accept incoming network connections. Incoming network connections are required to use parallel processing as implemented in multimarkClosed. Note that setting parms="all" is required for any multimarkClosed model output to be used in multimodelClosed .
Value
A list containing the following:
mcmc
Markov chain Monte Carlo object of class mcmc.list .
mod.p
Model formula for detection probability (as specified by mod.p above).
mod.delta
Model formula for conditional probability of type 1 or type 2 encounter, given detection (as specified by mod.delta above).
DM
A list of design matrices for detection probability generated for model mod.p, where DM$p is the design matrix for initial capture probability (p) and DM$c is the design matrix for recapture probability (c).
initial.values
A list containing the parameter and latent variable values at iteration iter for each chain. Values are provided for "pbeta", "N", "delta_1", "delta_2", "alpha", "sigma2_zp", "zp", "psi", "x", and "H".
mms
An object of class multimarksetup
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
McClintock, B. T., Bailey, L. L., Dreher, B. P., and Link, W. A. 2014. Probit models for capture-recapture data subject to imperfect detection, individual heterogeneity and misidentification. The Annals of Applied Statistics 8: 2461-2484.
See Also
bobcat , processdata , multimodelClosed
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Run single chain using the default model for bobcat data
bobcat.dot<-multimarkClosed(bobcat)
#Posterior summary for monitored parameters
summary(bobcat.dot$mcmc)
plot(bobcat.dot$mcmc)
Fit spatially-explicit population abundance models for capture-mark-recapture data consisting of multiple non-invasive marks
Description
This function fits spatially-explicit population abundance models for capture-mark-recapture data consisting of multiple non-invasive marks using Bayesian analysis methods. Markov chain Monte Carlo (MCMC) is used to draw samples from the joint posterior distribution.
Usage
multimarkClosedSCR(
Enc.Mat,
trapCoords,
studyArea = NULL,
buffer = NULL,
ncells = 1024,
data.type = "never",
covs = data.frame(),
mms = NULL,
mod.p = ~1,
mod.delta = ~type,
detection = "half-normal",
parms = c("pbeta", "delta", "N"),
nchains = 1,
iter = 12000,
adapt = 1000,
bin = 50,
thin = 1,
burnin = 2000,
taccept = 0.44,
tuneadjust = 0.95,
proppbeta = 0.1,
propsigma = 1,
propcenter = NULL,
maxnumbasis = 1,
a0delta = 1,
a0alpha = 1,
b0alpha = 1,
sigma_bounds = NULL,
mu0 = 0,
sigma2_mu0 = 1.75,
a0psi = 1,
b0psi = 1,
initial.values = NULL,
known = integer(),
scalemax = 10,
printlog = FALSE,
...
)
Arguments
Enc.Mat
A matrix containing the observed encounter histories with rows corresponding to individuals and (ntraps*noccas) columns corresponding to traps and sampling occasions. The first noccas columns correspond to trap 1, the second noccas columns corresopond to trap 2, etc. Ignored unless mms=NULL.
trapCoords
A matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate (“x”), and the second column the y-coordinate (“y”). The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating. Ignored unless mms=NULL.
studyArea
is a 3-column matrix containing the coordinates for the centroids of a contiguous grid of cells that define the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns (“x” and “y”) indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column (“avail”) indicates whether the cell is available habitat (=1) or not (=0). All cells must be square and have the same resolution. If studyArea=NULL (the default) and mms=NULL, then a square study area grid composed of ncells cells of available habitat is drawn around the bounding box of trapCoords based on buffer. Ignored unless mms=NULL.
Note that rows should be ordered in raster cell order (raster cell numbers start at 1 in the upper left corner, and increase from left to right, and then from top to bottom).
buffer
A scaler in same units as trapCoords indicating the buffer around the bounding box of trapCoords for defining the study area when studyArea=NULL. Ignored unless studyArea=NULL and mms=NULL.
ncells
The number of grid cells in the study area when studyArea=NULL. The square root of ncells must be a whole number. Default is ncells=1024. Ignored unless studyArea=NULL and mms=NULL.
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
covs
A data frame of time- and/or trap-dependent covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of traps times the number of sampling occasions (ntraps*noccas), where the first noccas rows correspond to trap 1, the second noccas rows correspond to trap 2, etc. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
mms
An optional object of class multimarkSCRsetup-class; if NULL it is created. See processdataSCR .
mod.p
Model formula for detection probability as a function of distance from activity centers. For example, mod.p=~1 specifies no effects (i.e., intercept only) other than distance, mod.p~time specifies temporal effects, mod.p~c specifies behavioral reponse (i.e., trap "happy" or "shy"), mod.p~trap specifies trap effects, and mod.p~time+c specifies additive temporal and behavioral effects.
mod.delta
Model formula for conditional probabilities of type 1 (delta_1) and type 2 (delta_2) encounters, given detection. Currently only mod.delta=~1 (i.e., \delta_1 = \delta_2) and mod.delta=~type (i.e., \delta_1 \ne \delta_2) are implemented.
detection
Model for detection probability as a function of distance from activity centers . Must be "half-normal" (of the form \exp{(-d^2 / (2*\sigma^2))}, where d is distance) or "exponential" (of the form \exp{(-d / \lambda)}).
parms
A character vector giving the names of the parameters and latent variables to monitor. Possible parameters are cloglog-scale detection probability parameters ("pbeta"), population abundance ("N"), conditional probability of type 1 or type 2 encounter, given detection ("delta)", probability of simultaneous type 1 and type 2 detection, given both types encountered ("alpha"), cloglog-scale distance term for the detection function ("sigma2_scr" when detection=``half-normal'' or "lambda" when detection=``exponential''), and the probability that a randomly selected individual from the M = nrow(Enc.Mat) observed individuals belongs to the n unique individuals encountered at least once ("psi"). Individual activity centers ("centers"), encounter history indices ("H"), and the log posterior density ("logPosterior") may also be monitored. Setting parms="all" monitors all possible parameters and latent variables.
nchains
The number of parallel MCMC chains for the model.
iter
The number of MCMC iterations.
adapt
The number of iterations for proposal distribution adaptation. If adapt = 0 then no adaptation occurs.
bin
Bin length for calculating acceptance rates during adaptive phase (0 < bin <= iter).
thin
Thinning interval for monitored parameters.
burnin
Number of burn-in iterations (0 <= burnin < iter).
taccept
Target acceptance rate during adaptive phase (0 < taccept <= 1). Acceptance rate is monitored every bin iterations. Default is taccept = 0.44.
tuneadjust
Adjustment term during adaptive phase (0 < tuneadjust <= 1). If acceptance rate is less than taccept, then proposal term (proppbeta or propsigma) is multiplied by tuneadjust. If acceptance rate is greater than or equal to taccept, then proposal term is divided by tuneadjust. Default is tuneadjust = 0.95.
proppbeta
Scaler or vector (of length k) specifying the initial standard deviation of the Normal(pbeta[j], proppbeta[j]) proposal distribution. If proppbeta is a scaler, then this value is used for all j = 1, ..., k. Default is proppbeta = 0.1.
propsigma
Scaler specifying the initial Gamma(shape = 1/propsigma, scale = sigma_scr * propsigma) proposal distribution for sigma_scr = sqrt(sigma2_scr) (or sqrt(lambda) = lambda if detection=``exponential''). Default is propsigma=1.
propcenter
Scaler specifying the neighborhood distance when proposing updates to activity centers. When propcenter=NULL (the default), then propcenter = a*10, where a is the cell size for the study area grid, and each cell has (at most) approximately 300 neighbors.
maxnumbasis
Maximum number of basis vectors to use when proposing latent history frequency updates. Default is maxnumbasis = 1, but higher values can potentially improve mixing.
a0delta
Scaler or vector (of length d) specifying the prior for the conditional (on detection) probability of type 1 (delta_1), type 2 (delta_2), and both type 1 and type 2 encounters (1-delta_1-delta_2). If a0delta is a scaler, then this value is used for all a0delta[j] for j = 1, ..., d. For mod.delta=~type, d=3 with [delta_1, delta_2, 1-delta_1-delta_2] ~ Dirichlet(a0delta) prior. For mod.delta=~1, d=2 with [tau] ~ Beta(a0delta[1],a0delta[2]) prior, where (delta_1,delta_2,1-delta_1-delta_2) = (tau/2,tau/2,1-tau). See McClintock et al. (2013) for more details.
a0alpha
Specifies "shape1" parameter for [alpha] ~ Beta(a0alpha, b0alpha) prior. Only applicable when data.type = "sometimes". Default is a0alpha = 1. Note that when a0alpha = 1 and b0alpha = 1, then [alpha] ~ Unif(0,1).
b0alpha
Specifies "shape2" parameter for [alpha] ~ Beta(a0alpha, b0alpha) prior. Only applicable when data.type = "sometimes". Default is b0alpha = 1. Note that when a0alpha = 1 and b0alpha = 1, then [alpha] ~ Unif(0,1).
sigma_bounds
Positive vector of length 2 for the lower and upper bounds for the [sigma_scr] ~ Uniform(sigma_bounds[1], sigma_bounds[2]) (or [sqrt(lambda)] when detection=``exponential'') prior for the detection function term sigma_scr = sqrt(sigma2_scr) (or sqrt(lambda)). When sigma_bounds = NULL (the default), then sigma_bounds = c(1.e-6,max(diff(range(studyArea[,"x"])),diff(range(studyArea[,"y"])))).
mu0
Scaler or vector (of length k) specifying mean of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is mu0 = 0.
sigma2_mu0
Scaler or vector (of length k) specifying variance of pbeta[j] ~ Normal(mu0[j], sigma2_mu0[j]) prior. If sigma2_mu0 is a scaler, then this value is used for all j = 1, ..., k. Default is sigma2_mu0 = 1.75.
a0psi
Specifies "shape1" parameter for [psi] ~ Beta(a0psi,b0psi) prior. Default is a0psi = 1.
b0psi
Specifies "shape2" parameter for [psi] ~ Beta(a0psi,b0psi) prior. Default is b0psi = 1.
initial.values
Optional list of nchain list(s) specifying initial values for parameters and latent variables. Default is initial.values = NULL, which causes initial values to be generated automatically. In addition to the parameters ("pbeta", "N", "delta_1", "delta_2", "alpha", "sigma2_scr", "centers", and "psi"), initial values can be specified for the initial latent history frequencies ("x") and initial individual encounter history indices ("H").
known
Optional integer vector indicating whether the encounter history of an individual is known with certainty (i.e., the observed encounter history is the true encounter history). Encounter histories with at least one type 4 encounter are automatically assumed to be known, and known does not need to be specified unless there exist encounter histories that do not contain a type 4 encounter that happen to be known with certainty (e.g., from independent telemetry studies). If specified, known = c(v_1,v_2,...,v_M) must be a vector of length M = nrow(Enc.Mat) where v_i = 1 if the encounter history for individual i is known (v_i = 0 otherwise). Note that known all-zero encounter histories (e.g., ‘000’) are ignored.
scalemax
Upper bound for internal re-scaling of grid cell centroid coordinates. Default is scalemax=10, which re-scales the centroids to be between 0 and 10. Re-scaling is done internally to avoid numerical overflows during model fitting. Ignored unless mms=NULL.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
...
Additional "parameters" arguments for specifying mod.p. See make.design.data .
Details
The first time multimarkSCRClosed is called, it will likely produce a firewall warning alerting users that R has requested the ability to accept incoming network connections. Incoming network connections are required to use parallel processing as implemented in multimarkClosed. Note that setting parms="all" is required for any multimarkClosed model output to be used in multimodelClosed .
Value
A list containing the following:
mcmc
Markov chain Monte Carlo object of class mcmc.list .
mod.p
Model formula for detection probability (as specified by mod.p above).
mod.delta
Model formula for conditional probability of type 1 or type 2 encounter, given detection (as specified by mod.delta above).
mod.det
Model formula for detection function (as specified by detection above).
DM
A list of design matrices for detection probability generated for model mod.p, where DM$p is the design matrix for initial capture probability (p) and DM$c is the design matrix for recapture probability (c).
initial.values
A list containing the parameter and latent variable values at iteration iter for each chain. Values are provided for "pbeta", "N", "delta_1", "delta_2", "alpha", "sigma2_scr", "centers", "psi", "x", and "H".
mms
An object of class multimarkSCRsetup
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
Gopalaswamy, A.M., Royle, J.A., Hines, J.E., Singh, P., Jathanna, D., Kumar, N. and Karanth, K.U. 2012. Program SPACECAP: software for estimating animal density using spatially explicit capture-recapture models. Methods in Ecology and Evolution 3:1067-1072.
King, R., McClintock, B. T., Kidney, D., and Borchers, D. L. 2016. Capture-recapture abundance estimation using a semi-complete data likelihood approach. The Annals of Applied Statistics 10: 264-285
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
McClintock, B. T., Bailey, L. L., Dreher, B. P., and Link, W. A. 2014. Probit models for capture-recapture data subject to imperfect detection, individual heterogeneity and misidentification. The Annals of Applied Statistics 8: 2461-2484.
Royle, J.A., Karanth, K.U., Gopalaswamy, A.M. and Kumar, N.S. 2009. Bayesian inference in camera trapping studies for a class of spatial capture-recapture models. Ecology 90: 3233-3244.
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Generate object of class "multimarkSCRsetup" from simulated data
sim.data<-simdataClosedSCR()
Enc.Mat <- sim.data$Enc.Mat
trapCoords <- sim.data$spatialInputs$trapCoords
studyArea <- sim.data$spatialInputs$studyArea
#Run single chain using the default model for simulated data
example.dot<-multimarkClosedSCR(Enc.Mat,trapCoords,studyArea)
#Posterior summary for monitored parameters
summary(example.dot$mcmc)
plot(example.dot$mcmc)
Class "multimarkSCRsetup"
Description
A class of spatial 'mulitmark' model inputs
Slots
Enc.MatObject of class
"matrix". The observed encounter histories (with rows corresponding to individuals and columns corresponding to sampling occasions).data.typeObject of class
"character". The encounter history data type ("never", "sometimes", or "always").vAll.histsObject of class
"integer". An ordered vector containing all possible encounter histories in sequence.AprimeObject of class
"sparseMatrix". Transpose of the A matrix mapping latent encounter histories to observed histories.indBasisObject of class
"numeric".An ordered vector of the indices of the three encounter histories updated by each basis vector.ncolbasisObject of class
"integer". The number of needed basis vectors.knownxObject of class
"integer". Frequencies of known encounter histories.CObject of class
"integer". Sampling occasion of first capture for each encounter history.LObject of class
"integer". Sampling occasion of last capture for each encounter history.naivexObject of class
"integer". “Naive” latent history frequencies assuming a one-to-one mapping withEnc.Mat.covsObject of class
"data.frame". Temporal covariates for detection probability (the number of rows in the data frame must equal the number of sampling occasions).spatialInputsObject of class
"list". List is of length 4 containingtrapCoordsandstudyAreaafter re-scaling coordinates based onmaxscale, as well as the original (not re-scaled) grid cell resolution (origCellRes) and re-scaling range (Srange).
Objects from the Class
Objects can be created by calls of the form processdata(Enc.Mat, ...) or new("multimarkSCRsetup", ...).
Methods
No methods defined with class "multimarkSCRsetup".
Author(s)
Brett T. McClintock
See Also
Examples
showClass("multimarkSCRsetup")
Class "multimarksetup"
Description
A class of 'mulitmark' model inputs
Slots
Enc.MatObject of class
"matrix". The observed encounter histories (with rows corresponding to individuals and columns corresponding to sampling occasions).data.typeObject of class
"character". The encounter history data type ("never", "sometimes", or "always").vAll.histsObject of class
"integer". An ordered vector containing all possible encounter histories in sequence.AprimeObject of class
"sparseMatrix". Transpose of the A matrix mapping latent encounter histories to observed histories.indBasisObject of class
"numeric".An ordered vector of the indices of the three encounter histories updated by each basis vector.ncolbasisObject of class
"integer". The number of needed basis vectors.knownxObject of class
"integer". Frequencies of known encounter histories.CObject of class
"integer". Sampling occasion of first capture for each encounter history.LObject of class
"integer". Sampling occasion of last capture for each encounter history.naivexObject of class
"integer". “Naive” latent history frequencies assuming a one-to-one mapping withEnc.Mat.covsObject of class
"data.frame". Temporal covariates for detection probability (the number of rows in the data frame must equal the number of sampling occasions).
Objects from the Class
Objects can be created by calls of the form processdata(Enc.Mat, ...) or new("multimarksetup", ...).
Methods
No methods defined with class "multimarksetup".
Author(s)
Brett T. McClintock
See Also
Examples
showClass("multimarksetup")
Multimodel inference for 'multimark' open population survival models
Description
This function performs Bayesian multimodel inference for a set of 'multimark' open population survival (i.e., Cormack-Jolly-Seber) models using the reversible jump Markov chain Monte Carlo (RJMCMC) algorithm proposed by Barker & Link (2013).
Usage
multimodelCJS(
modlist,
modprior = rep(1/length(modlist), length(modlist)),
monparms = "phi",
miter = NULL,
mburnin = 0,
mthin = 1,
M1 = NULL,
pbetapropsd = 1,
zppropsd = NULL,
phibetapropsd = 1,
zphipropsd = NULL,
sigppropshape = 1,
sigppropscale = 0.01,
sigphipropshape = 1,
sigphipropscale = 0.01,
printlog = FALSE
)
Arguments
modlist
A list of individual model output lists returned by multimarkCJS . The models must have the same number of chains and MCMC iterations.
modprior
Vector of length length(modlist) containing prior model probabilities. Default is modprior = rep(1/length(modlist), length(modlist)).
monparms
Parameters to monitor. Only parameters common to all models can be monitored (e.g., "pbeta[(Intercept)]", "phibeta[(Intercept)]", "psi"), but derived survival ("phi") and capture ("p") probabilities can also be monitored. Default is monparms = "phi".
miter
The number of RJMCMC iterations per chain. If NULL, then the number of MCMC iterations for each individual model chain is used.
mburnin
Number of burn-in iterations (0 <= mburnin < miter).
mthin
Thinning interval for monitored parameters.
M1
Integer vector indicating the initial model for each chain, where M1_j=i initializes the RJMCMC algorithm for chain j in the model corresponding to modlist[[i]] for i=1,..., length(modlist). If NULL, the algorithm for all chains is initialized in the most general model. Default is M1=NULL.
pbetapropsd
Scaler specifying the standard deviation of the Normal(0, pbetapropsd) proposal distribution for "pbeta" parameters. Default is pbetapropsd=1. See Barker & Link (2013) for more details.
zppropsd
Scaler specifying the standard deviation of the Normal(0, zppropsd) proposal distribution for "zp" parameters. Only applies if at least one (but not all) model(s) include individual hetergeneity in detection probability. If NULL, zppropsd = sqrt(sigma2_zp) is used. Default is zppropsd=NULL. See Barker & Link (2013) for more details.
phibetapropsd
Scaler specifying the standard deviation of the Normal(0, phibetapropsd) proposal distribution for "phibeta" parameters. Default is phibetapropsd=1. See Barker & Link (2013) for more details.
zphipropsd
Scaler specifying the standard deviation of the Normal(0, zphipropsd) proposal distribution for "zphi" parameters. Only applies if at least one (but not all) model(s) include individual hetergeneity in survival probability. If NULL, zphipropsd = sqrt(sigma2_zphi) is used. Default is zphipropsd=NULL. See Barker & Link (2013) for more details.
sigppropshape
Scaler specifying the shape parameter of the invGamma(shape = sigppropshape, scale = sigppropscale) proposal distribution for "sigma2_zp". Only applies if at least one (but not all) model(s) include individual hetergeneity in detection probability. Default is sigppropshape=1. See Barker & Link (2013) for more details.
sigppropscale
Scaler specifying the scale parameter of the invGamma(shape = sigppropshape, scale = sigppropscale) proposal distribution for "sigma2_zp". Only applies if at least one (but not all) model(s) include individual hetergeneity in detection probability. Default is sigppropscale=0.01. See Barker & Link (2013) for more details.
sigphipropshape
Scaler specifying the shape parameter of the invGamma(shape = sigphipropshape, scale = sigphipropscale) proposal distribution for "sigma2_zphi". Only applies if at least one (but not all) model(s) include individual hetergeneity in survival probability. Default is sigphipropshape=1. See Barker & Link (2013) for more details.
sigphipropscale
Scaler specifying the scale parameter of the invGamma(shape = sigphipropshape, scale = sigphipropscale) proposal distribution for "sigma_zphi". Only applies if at least one (but not all) model(s) include individual hetergeneity in survival probability. Default is sigphipropscale=0.01. See Barker & Link (2013) for more details.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
Details
Note that setting parms="all" is required when fitting individual multimarkCJS models to be included in modlist.
Value
A list containing the following:
rjmcmc
Reversible jump Markov chain Monte Carlo object of class mcmc.list . Includes RJMCMC output for monitored parameters and the current model at each iteration ("M").
pos.prob
A list of calculated posterior model probabilities for each chain, including the overall posterior model probabilities across all chains.
Author(s)
Brett T. McClintock
References
Barker, R. J. and Link. W. A. 2013. Bayesian multimodel inference by RJMCMC: a Gibbs sampling approach. The American Statistician 67: 150-156.
See Also
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Generate object of class "multimarksetup" from simulated data
data_type = "always"
noccas <- 5
phibetaTime <- seq(2,0,length=noccas-1) # declining trend in survival
data <- simdataCJS(noccas=5,phibeta=phibetaTime,data.type=data_type)
setup <- processdata(data$Enc.Mat,data.type=data_type)
#Run single chain using the default model. Note parms="all".
sim.pdot.phidot <- multimarkCJS(mms=setup,parms="all",iter=1000,adapt=500,burnin=500)
#Run single chain with temporal trend for phi. Note parms="all".
sim.pdot.phiTime <- multimarkCJS(mms=setup,mod.phi=~Time,parms="all",iter=1000,adapt=500,burnin=500)
#Perform RJMCMC using defaults
modlist <- list(mod1=sim.pdot.phidot,mod2=sim.pdot.phiTime)
sim.M <- multimodelCJS(modlist=modlist)
#Posterior model probabilities
sim.M$pos.prob
#multimodel posterior summary for survival (display first cohort only)
summary(sim.M$rjmcmc[,paste0("phi[1,",1:(noccas-1),"]")])
Multimodel inference for 'multimark' closed population abundance models
Description
This function performs Bayesian multimodel inference for a set of 'multimark' closed population abundance models using the reversible jump Markov chain Monte Carlo (RJMCMC) algorithm proposed by Barker & Link (2013).
Usage
multimodelClosed(
modlist,
modprior = rep(1/length(modlist), length(modlist)),
monparms = "N",
miter = NULL,
mburnin = 0,
mthin = 1,
M1 = NULL,
pbetapropsd = 1,
zppropsd = NULL,
sigppropshape = 6,
sigppropscale = 4,
printlog = FALSE
)
Arguments
modlist
A list of individual model output lists returned by multimarkClosed or markClosed . The models must have the same number of chains and MCMC iterations.
modprior
Vector of length length(modlist) containing prior model probabilities. Default is modprior = rep(1/length(modlist), length(modlist)).
monparms
Parameters to monitor. Only parameters common to all models can be monitored (e.g., "pbeta[(Intercept)]", "N"), but derived capture ("p") and recapture ("c") probabilities can also be monitored. Default is monparms = "N".
miter
The number of RJMCMC iterations per chain. If NULL, then the number of MCMC iterations for each individual model chain is used.
mburnin
Number of burn-in iterations (0 <= mburnin < miter).
mthin
Thinning interval for monitored parameters.
M1
Integer vector indicating the initial model for each chain, where M1_j=i initializes the RJMCMC algorithm for chain j in the model corresponding to modlist[[i]] for i=1,..., length(modlist). If NULL, the algorithm for all chains is initialized in the most general model. Default is M1=NULL.
pbetapropsd
Scaler specifying the standard deviation of the Normal(0, pbetapropsd) proposal distribution for "pbeta" parameters. Default is pbetapropsd=1. See Barker & Link (2013) for more details.
zppropsd
Scaler specifying the standard deviation of the Normal(0, zppropsd) proposal distribution for "zp" parameters. Only applies if at least one (but not all) model(s) include individual hetergeneity in detection probability. If NULL, zppropsd = sqrt(sigma2_zp) is used. Default is zppropsd=NULL. See Barker & Link (2013) for more details.
sigppropshape
Scaler specifying the shape parameter of the invGamma(shape = sigppropshape, scale = sigppropscale) proposal distribution for sigma_zp. Only applies if at least one (but not all) model(s) include individual hetergeneity in detection probability. Default is sigppropshape=6. See Barker & Link (2013) for more details.
sigppropscale
Scaler specifying the scale parameter of the invGamma(shape = sigppropshape, scale = sigppropscale) proposal distribution for sigma_zp. Only applies if at least one (but not all) model(s) include individual hetergeneity in detection probability. Default is sigppropscale=4. See Barker & Link (2013) for more details.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
Details
Note that setting parms="all" is required when fitting individual multimarkClosed or markClosed models to be included in modlist.
Value
A list containing the following:
rjmcmc
Reversible jump Markov chain Monte Carlo object of class mcmc.list . Includes RJMCMC output for monitored parameters and the current model at each iteration ("M").
pos.prob
A list of calculated posterior model probabilities for each chain, including the overall posterior model probabilities across all chains.
Author(s)
Brett T. McClintock
References
Barker, R. J. and Link. W. A. 2013. Bayesian multimodel inference by RJMCMC: a Gibbs sampling approach. The American Statistician 67: 150-156.
See Also
multimarkClosed , markClosed , processdata
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Generate object of class "multimarksetup"
setup <- processdata(bobcat)
#Run single chain using the default model for bobcat data. Note parms="all".
bobcat.dot <- multimarkClosed(mms=setup,parms="all",iter=1000,adapt=500,burnin=500)
#Run single chain for bobcat data with time effects. Note parms="all".
bobcat.time <- multimarkClosed(mms=setup,mod.p=~time,parms="all",iter=1000,adapt=500,burnin=500)
#Perform RJMCMC using defaults
modlist <- list(mod1=bobcat.dot,mod2=bobcat.time)
bobcat.M <- multimodelClosed(modlist=modlist,monparms=c("N","p"))
#Posterior model probabilities
bobcat.M$pos.prob
#multimodel posterior summary for abundance
summary(bobcat.M$rjmcmc[,"N"])
Multimodel inference for 'multimark' spatial population abundance models
Description
This function performs Bayesian multimodel inference for a set of 'multimark' spatial population abundance models using the reversible jump Markov chain Monte Carlo (RJMCMC) algorithm proposed by Barker & Link (2013).
Usage
multimodelClosedSCR(
modlist,
modprior = rep(1/length(modlist), length(modlist)),
monparms = "N",
miter = NULL,
mburnin = 0,
mthin = 1,
M1 = NULL,
pbetapropsd = 1,
sigpropmean = 0.8,
sigpropsd = 0.4,
printlog = FALSE
)
Arguments
modlist
A list of individual model output lists returned by multimarkClosedSCR or markClosedSCR . The models must have the same number of chains and MCMC iterations.
modprior
Vector of length length(modlist) containing prior model probabilities. Default is modprior = rep(1/length(modlist), length(modlist)).
monparms
Parameters to monitor. Only parameters common to all models can be monitored (e.g., "pbeta[(Intercept)]", "N", "sigma2_scr"), but derived density ("D") as well as capture ("p") and recapture ("c") probabilities (at distance zero from activity centers) can also be monitored. Default is monparms = "N".
miter
The number of RJMCMC iterations per chain. If NULL, then the number of MCMC iterations for each individual model chain is used.
mburnin
Number of burn-in iterations (0 <= mburnin < miter).
mthin
Thinning interval for monitored parameters.
M1
Integer vector indicating the initial model for each chain, where M1_j=i initializes the RJMCMC algorithm for chain j in the model corresponding to modlist[[i]] for i=1,..., length(modlist). If NULL, the algorithm for all chains is initialized in the most general model. Default is M1=NULL.
pbetapropsd
Scaler specifying the standard deviation of the Normal(0, pbetapropsd) proposal distribution for "pbeta" parameters. Default is pbetapropsd=1. See Barker & Link (2013) for more details.
sigpropmean
Scaler specifying the mean of the inverse Gamma proposal distribution for sigma2_scr (or lambda if detection=``exponential''). Only applies if models do not have the same detection function (i.e., “half-normal” or “exponential”). Default is sigpropmean=0.8. See Barker & Link (2013) for more details.
sigpropsd
Scaler specifying the standard deviation of the inverse Gamma proposal distribution for sigma2_scr (or lambda if detection=``exponential''). Only applies if models do not have the same detection function (i.e., “half-normal” or “exponential”). Default is sigpropsd=0.4. See Barker & Link (2013) for more details.
printlog
Logical indicating whether to print the progress of chains and any errors to a log file in the working directory. Ignored when nchains=1. Updates are printed to log file as 1% increments of iter of each chain are completed. With >1 chains, setting printlog=TRUE is probably most useful for Windows users because progress and errors are automatically printed to the R console for "Unix-like" machines (i.e., Mac and Linux) when printlog=FALSE. Default is printlog=FALSE.
Details
Note that setting parms="all" is required when fitting individual multimarkClosedSCR or markClosedSCR models to be included in modlist.
Value
A list containing the following:
rjmcmc
Reversible jump Markov chain Monte Carlo object of class mcmc.list . Includes RJMCMC output for monitored parameters and the current model at each iteration ("M").
pos.prob
A list of calculated posterior model probabilities for each chain, including the overall posterior model probabilities across all chains.
Author(s)
Brett T. McClintock
References
Barker, R. J. and Link. W. A. 2013. Bayesian multimodel inference by RJMCMC: a Gibbs sampling approach. The American Statistician 67: 150-156.
See Also
multimarkClosedSCR , processdataSCR
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Generate object of class "multimarkSCRsetup"
sim.data<-simdataClosedSCR()
Enc.Mat<-sim.data$Enc.Mat
trapCoords<-sim.data$spatialInputs$trapCoords
studyArea<-sim.data$spatialInputs$studyArea
setup<-processdataSCR(Enc.Mat,trapCoords,studyArea)
#Run single chain using the default model for simulated data. Note parms="all".
example.dot <- multimarkClosedSCR(mms=setup,parms="all",iter=1000,adapt=500,burnin=500)
#Run single chain for simulated data with behavior effects. Note parms="all".
example.c <- multimarkClosedSCR(mms=setup,mod.p=~c,parms="all",iter=1000,adapt=500,burnin=500)
#Perform RJMCMC using defaults
modlist <- list(mod1=example.dot,mod2=example.c)
example.M <- multimodelClosedSCR(modlist=modlist,monparms=c("N","D","sigma2_scr"))
#Posterior model probabilities
example.M$pos.prob
#multimodel posterior summary for abundance and density
summary(example.M$rjmcmc[,c("N","D")])
Plot spatial capture-mark-recapture data
Description
This function plots the study area grid, available habitat, and trap coordinates for spatial capture-recapture studies. Activity centers and capture locations can also be plotted.
Usage
plotSpatialData(
mms = NULL,
trapCoords,
studyArea,
centers = NULL,
trapLines = FALSE
)
Arguments
mms
An optional object of class multimarkSCRsetup-class from which the (re-scaled) study area and trap coordinates are plotted.
trapCoords
A matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate, and the second column the y-coordinate. The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating. Ignored unless mms=NULL.
studyArea
A 3-column matrix defining the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column indicates whether the cell is available habitat (=1) or not (=0). All cells must have the same resolution. Ignored unless mms=NULL.
Note that rows should be ordered in raster cell order (raster cell numbers start at 1 in the upper left corner, and increase from left to right, and then from top to bottom).
centers
An optional vector indicating the grid cell (i.e., the row of studyArea) that contains the true (latent) activity centers for each individual. If mms is provided, then centers must be of length nrow(Enc.Mat) (i.e., a center must be provided for each observed individual).
trapLines
Logical indicating whether to draw lines from activity centers to respective traps at which each individual was captured. Default is trapLines=FALSE. Ignored when mms=NULL or centers=NULL.
Author(s)
Brett T. McClintock
Examples
#Plot the tiger example data
plotSpatialData(trapCoords=tiger$trapCoords,studyArea=tiger$studyArea)
Generate model inputs for fitting 'multimark' models
Description
This function generates an object of class multimarksetup that is required to fit ‘multimark’ models.
Usage
processdata(
Enc.Mat,
data.type = "never",
covs = data.frame(),
known = integer()
)
Arguments
Enc.Mat
A matrix of observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions (ignored unless mms=NULL).
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
covs
A data frame of temporal covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of sampling occasions. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
known
Optional integer vector indicating whether the encounter history of an individual is known with certainty (i.e., the observed encounter history is the true encounter history). Encounter histories with at least one type 4 encounter are automatically assumed to be known, and known does not need to be specified unless there exist encounter histories that do not contain a type 4 encounter that happen to be known with certainty (e.g., from independent telemetry studies). If specified, known = c(v_1,v_2,...,v_M) must be a vector of length M = nrow(Enc.Mat) where v_i = 1 if the encounter history for individual i is known (v_i = 0 otherwise). Note that known all-zero encounter histories (e.g., ‘000’) are ignored.
Value
An object of class multimarksetup.
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
See Also
multimarksetup-class , multimarkClosed , bobcat
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Generate object of class "multimarksetup"
setup <- processdata(bobcat)
#Run single chain using the default model for bobcat data
bobcat.dot<-multimarkClosed(mms=setup)
#Run single chain for bobcat data with temporal effects (i.e., mod.p=~time)
bobcat.time <- multimarkClosed(mms=setup,mod.p=~time)
Generate model inputs for fitting spatial 'multimark' models
Description
This function generates an object of class multimarkSCRsetup that is required to fit spatial ‘multimark’ models.
Usage
processdataSCR(
Enc.Mat,
trapCoords,
studyArea = NULL,
buffer = NULL,
ncells = NULL,
data.type = "never",
covs = data.frame(),
known = integer(),
scalemax = 10
)
Arguments
Enc.Mat
A matrix containing the observed encounter histories with rows corresponding to individuals and (ntraps*noccas) columns corresponding to traps and sampling occasions. The first noccas columns correspond to trap 1, the second noccas columns corresopond to trap 2, etc. Ignored unless mms=NULL.
trapCoords
A matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate, and the second column the y-coordinate. The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating.
studyArea
is a 3-column matrix containing the coordinates for the centroids of a contiguous grid of cells that define the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column indicates whether the cell is available habitat (=1) or not (=0). All cells must be square and have the same resolution. If studyArea=NULL (the default), then a square study area grid composed of ncells cells of available habitat is drawn around the bounding box of trapCoords based on buffer.
Note that rows should be ordered in raster cell order (raster cell numbers start at 1 in the upper left corner, and increase from left to right, and then from top to bottom).
buffer
A scaler in same units as trapCoords indicating the buffer around the bounding box of trapCoords for defining the study area when studyArea=NULL. Ignored unless studyArea=NULL.
ncells
The number of grid cells in the study area when studyArea=NULL. The square root of ncells must be a whole number. Default is ncells=1024. Ignored unless studyArea=NULL.
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
covs
A data frame of time- and/or trap-dependent covariates for detection probabilities (ignored unless mms=NULL). The number of rows in the data frame must equal the number of traps times the number of sampling occasions (ntraps*noccas), where the first noccas rows correspond to trap 1, the second noccas rows correspond to trap 2, etc. Covariate names cannot be "time", "age", or "h"; these names are reserved for temporal, behavioral, and individual effects when specifying mod.p and mod.phi.
known
Optional integer vector indicating whether the encounter history of an individual is known with certainty (i.e., the observed encounter history is the true encounter history). Encounter histories with at least one type 4 encounter are automatically assumed to be known, and known does not need to be specified unless there exist encounter histories that do not contain a type 4 encounter that happen to be known with certainty (e.g., from independent telemetry studies). If specified, known = c(v_1,v_2,...,v_M) must be a vector of length M = nrow(Enc.Mat) where v_i = 1 if the encounter history for individual i is known (v_i = 0 otherwise). Note that known all-zero encounter histories (e.g., ‘000’) are ignored.
scalemax
Upper bound for internal re-scaling of grid cell centroid coordinates. Default is scalemax=10, which re-scales the centroids to be between 0 and 10. Re-scaling is done internally to avoid numerical overflows during model fitting.
Value
An object of class multimarkSCRsetup.
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
Gopalaswamy, A.M., Royle, J.A., Hines, J.E., Singh, P., Jathanna, D., Kumar, N. and Karanth, K.U. 2012. Program SPACECAP: software for estimating animal density using spatially explicit capture-recapture models. Methods in Ecology and Evolution 3:1067-1072.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
Royle, J.A., Karanth, K.U., Gopalaswamy, A.M. and Kumar, N.S. 2009. Bayesian inference in camera trapping studies for a class of spatial capture-recapture models. Ecology 90: 3233-3244.
See Also
multimarkSCRsetup-class , multimarkClosedSCR
Examples
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
#Generate object of class "multimarksetup" from simulated data
sim.data<-simdataClosedSCR()
Enc.Mat <- sim.data$Enc.Mat
trapCoords <- sim.data$spatialInputs$trapCoords
studyArea <- sim.data$spatialInputs$studyArea
setup <- processdataSCR(Enc.Mat,trapCoords,studyArea)
#Run single chain using the default model for simulated data
example.dot<-multimarkClosedSCR(mms=setup)
Simulate open population capture-mark-recapture data arising from multiple non-invasive marks
Description
This function generates encounter histories from simulated open population capture-mark-recapture data consisting of multiple non-invasive marks.
Usage
simdataCJS(
N = 100,
noccas = 5,
pbeta = -0.25,
sigma2_zp = 0,
phibeta = 1.6,
sigma2_zphi = 0,
delta_1 = 0.4,
delta_2 = 0.4,
alpha = 0.5,
data.type = "never",
link = "probit"
)
Arguments
N
Number of individuals.
noccas
Number of sampling occasions. floor(N/noccas) individuals are first encountered on each occasion.
pbeta
Logit- or probit-scale intercept term(s) for capture probability (p). Must be a scaler or vector of length noccas.
sigma2_zp
Logit- or probit-scale individual heterogeneity variance term for capture probability (p).
phibeta
Logit- or probit-scale intercept term(s) for survival probability (\phi). Must be a scaler or vector of length noccas.
sigma2_zphi
Logit- or probit-scale individual heterogeneity variance term for survival probability (\phi).
delta_1
Conditional probability of type 1 encounter, given detection.
delta_2
Conditional probability of type 2 encounter, given detection.
alpha
Conditional probability of simultaneous type 1 and type 2 detection, given both types encountered. Only applies when data.type="sometimes".
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
link
Link function for detection probability. Must be "logit" or "probit". Note that multimarkCJS is currently implemented for the probit link only.
Value
A list containing the following:
Enc.Mat
A matrix containing the observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions.
trueEnc.Mat
A matrix containing the true (latent) encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions.
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
See Also
Examples
#simulate data for data.type="sometimes" using defaults
data<-simdataCJS(data.type="sometimes")
Simulate closed population capture-mark-recapture data arising from multiple non-invasive marks
Description
This function generates encounter histories from simulated closed population capture-mark-recapture data consisting of multiple non-invasive marks.
Usage
simdataClosed(
N = 100,
noccas = 5,
pbeta = -0.4,
tau = 0,
sigma2_zp = 0,
delta_1 = 0.4,
delta_2 = 0.4,
alpha = 0.5,
data.type = "never",
link = "logit"
)
Arguments
N
True population size or abundance.
noccas
The number of sampling occasions.
pbeta
Logit- or probit-scale intercept term(s) for capture probability (p). Must be a scaler or vector of length noccas.
tau
Additive logit- or probit-scale behavioral effect term for recapture probability (c).
sigma2_zp
Logit- or probit-scale individual heterogeneity variance term.
delta_1
Conditional probability of type 1 encounter, given detection.
delta_2
Conditional probability of type 2 encounter, given detection.
alpha
Conditional probability of simultaneous type 1 and type 2 detection, given both types encountered. Only applies when data.type="sometimes".
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
link
Link function for detection probability. Must be "logit" or "probit". Note that multimarkClosed is currently implemented for the logit link only.
Value
A list containing the following:
Enc.Mat
A matrix containing the observed encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions.
trueEnc.Mat
A matrix containing the true (latent) encounter histories with rows corresponding to individuals and columns corresponding to sampling occasions.
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
See Also
Examples
#simulate data for data.type="sometimes" using defaults
data<-simdataClosed(data.type="sometimes")
Simulate spatially-explicit capture-mark-recapture data from a (demographically) closed population with multiple non-invasive marks
Description
This function generates encounter histories from spatially-explicit capture-mark-recapture data consisting of multiple non-invasive marks.
Usage
simdataClosedSCR(
N = 30,
ntraps = 9,
noccas = 5,
pbeta = 0.25,
tau = 0,
sigma2_scr = 0.75,
lambda = 0.75,
delta_1 = 0.4,
delta_2 = 0.4,
alpha = 0.5,
data.type = "never",
detection = "half-normal",
spatialInputs = NULL,
buffer = 3 * sqrt(sigma2_scr),
ncells = 1024,
scalemax = 10,
plot = TRUE
)
Arguments
N
True population size or abundance.
ntraps
The number of traps. If trapCoords=NULL, the square root of ntraps must be a whole number in order to create a regular grid of trap coordinates on a square.
noccas
Scaler indicating the number of sampling occasions per trap.
pbeta
Complementary loglog-scale intercept term for detection probability (p). Must be a scaler or vector of length noccas.
tau
Additive complementary loglog-scale behavioral effect term for recapture probability (c).
sigma2_scr
Complementary loglog-scale term for effect of distance in the “half-normal” detection function. Ignored unless detection=``half-normal''.
lambda
Complementary loglog-scale term for effect of distance in the “exponential” detection function. Ignored unless detection=``exponential''.
delta_1
Conditional probability of type 1 encounter, given detection.
delta_2
Conditional probability of type 2 encounter, given detection.
alpha
Conditional probability of simultaneous type 1 and type 2 detection, given both types encountered. Only applies when data.type="sometimes".
data.type
Specifies the encounter history data type. All data types include non-detections (type 0 encounter), type 1 encounter (e.g., left-side), and type 2 encounters (e.g., right-side). When both type 1 and type 2 encounters occur for the same individual within a sampling occasion, these can either be "non-simultaneous" (type 3 encounter) or "simultaneous" (type 4 encounter). Three data types are currently permitted:
data.type="never" indicates both type 1 and type 2 encounters are never observed for the same individual within a sampling occasion, and observed encounter histories therefore include only type 1 or type 2 encounters (e.g., only left- and right-sided photographs were collected). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), and type 2 encounters (2). See bobcat . Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 3 encounters (3).
data.type="sometimes" indicates both type 1 and type 2 encounters are sometimes observed (e.g., both-sided photographs are sometimes obtained, but not necessarily for all individuals). Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4). Type 3 encounters can only be observed when an individual has at least one type 4 encounter. Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), type 3 encounters (3), and type 4 encounters (4).
data.type="always" indicates both type 1 and type 2 encounters are always observed, but some encounter histories may still include only type 1 or type 2 encounters. Observed encounter histories can consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4). Latent encounter histories consist of non-detections (0), type 1 encounters (1), type 2 encounters (2), and type 4 encounters (4).
detection
Model for detection probability as a function of distance from activity centers. Must be "half-normal" (of the form \exp{(-d^2 / (2*\sigma^2))}, where d is distance) or "exponential" (of the form \exp{(-d / \lambda)}).
spatialInputs
A list of length 3 composed of objects named trapCoords, studyArea, and centers:
trapCoords is a matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate (“x”), and the second column the y-coordinate (“y”). The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating.
studyArea is a 3-column matrix defining the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns (“x” and “y”) indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column (“avail”) indicates whether the cell is available habitat (=1) or not (=0). All grid cells must have the same resolution. Note that rows should be ordered in raster cell order (raster cell numbers start at 1 in the upper left corner, and increase from left to right, and then from top to bottom).
centers is a N-vector indicating the grid cell (i.e., the row of studyArea) that contains the true (latent) activity centers for each individual in the population.
If spatialInputs=NULL (the default), then all traps are assumed to be operating on all occasions, the study area is assumed to be composed of ncells grid cells, grid cells within buffer of the trap array are assumed to be available habitat, and the activity centers are randomly assigned to grid cells of available habitat.
buffer
A scaler indicating the buffer around the bounding box of trapCoords for defining the study area and available habitat when spatialInputs=NULL. Default is buffer=3*sqrt(sigma2_scr). Ignored unless spatialInputs=NULL.
ncells
The number of grid cells in the study area when studyArea=NULL. The square root of ncells must be a whole number. Default is ncells=1024. Ignored unless spatialInputs=NULL.
scalemax
Upper bound for grid cell centroid x- and y-coordinates. Default is scalemax=10, which scales the x- and y-coordinates to be between 0 and 10. Ignored unless spatialInputs=NULL.
plot
Logical indicating whether to plot the simulated trap coordinates, study area, and activity centers using plotSpatialData . Default is plot=TRUE
Details
Please be very careful when specifying your own spatialInputs; multimarkClosedSCR and markClosedSCR do little to verify that these make sense during model fitting.
Value
A list containing the following:
Enc.Mat
Matrix containing the observed encounter histories with rows corresponding to individuals and (ntraps*noccas) columns corresponding to traps and sampling occasions. The first noccas columns correspond to trap 1, the second noccas columns corresopond to trap 2, etc.
trueEnc.Mat
Matrix containing the true (latent) encounter histories with rows corresponding to individuals and (ntraps*noccas) columns corresponding to traps and sampling occasions. The first noccas columns correspond to trap 1, the second noccas columns corresopond to trap 2, etc.
spatialInputs
List of length 2 with objects named trapCoords and studyArea:
trapCoords is a matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate, and the second column the y-coordinate. The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating.
studyArea is a 3-column matrix containing the coordinates for the centroids a contiguous grid of cells that define the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column indicates whether the cell is available habitat (=1) or not (=0). All cells must have the same resolution.
centers
N-vector indicating the grid cell (i.e., the row of spatialInputs$studyArea) that contains the true (latent) activity centers for each individual in the population.
Author(s)
Brett T. McClintock
References
Bonner, S. J., and Holmberg J. 2013. Mark-recapture with multiple, non-invasive marks. Biometrics 69: 766-775.
McClintock, B. T., Conn, P. B., Alonso, R. S., and Crooks, K. R. 2013. Integrated modeling of bilateral photo-identification data in mark-recapture analyses. Ecology 94: 1464-1471.
Royle, J.A., Karanth, K.U., Gopalaswamy, A.M. and Kumar, N.S. 2009. Bayesian inference in camera trapping studies for a class of spatial capture-recapture models. Ecology 90: 3233-3244.
See Also
processdataSCR , multimarkClosedSCR , markClosedSCR
Examples
#simulate data for data.type="sometimes" using defaults
data<-simdataClosedSCR(data.type="sometimes")
Tiger data
Description
Example tiger data for multimark package.
Format
These spatial capture-recapture data with a single mark type are summarized in a list of length 3 containing the following objects:
Enc.Mat is a 44 x (noccas*ntraps) matrix containing observed encounter histories for 44 tigers across noccas=48 sampling occasions and ntraps=120 traps.
trapCoords is a matrix of dimension ntraps x (2 + noccas) indicating the Cartesian coordinates and operating occasions for the traps, where rows correspond to trap, the first column the x-coordinate, and the second column the y-coordinate. The last noccas columns indicate whether or not the trap was operating on each of the occasions, where ‘1’ indciates the trap was operating and ‘0’ indicates the trap was not operating.
studyArea is a 3-column matrix containing the coordinates for the centroids of the contiguous grid of cells that define the study area and available habitat. Each row corresponds to a grid cell. The first 2 columns indicate the Cartesian x- and y-coordinate for the centroid of each grid cell, and the third column indicates whether the cell is available habitat (=1) or not (=0). The grid cells are 0.336 km^2 resolution.
These data were obtained from the R package SPACECAP and modified by projecting onto a regular rectangular grid consisting of square grid cells (as is required by the spatial capture-recapture models in multimark).
Details
We thank Ullas Karanth, Wildlife Conservation Society, for providing the tiger data for use as an example with this package.
Source
Gopalaswamy, A.M., Royle, J.A., Hines, J.E., Singh, P., Jathanna, D., Kumar, N. and Karanth, K.U. 2012. Program SPACECAP: software for estimating animal density using spatially explicit capture-recapture models. Methods in Ecology and Evolution 3:1067-1072.
Royle, J.A., Karanth, K.U., Gopalaswamy, A.M. and Kumar, N.S. 2009. Bayesian inference in camera trapping studies for a class of spatial capture-recapture models. Ecology 90: 3233-3244.
See Also
Examples
data(tiger)
#plot the traps and available habitat within the study area
plotSpatialData(trapCoords=tiger$trapCoords,studyArea=tiger$studyArea)
# This example is excluded from testing to reduce package check time
# Example uses unrealistically low values for nchain, iter, and burnin
# Fit spatial model to tiger data
Enc.Mat<-tiger$Enc.Mat
trapCoords<-tiger$trapCoords
studyArea<-tiger$studyArea
tiger.dot<-markClosedSCR(Enc.Mat,trapCoords,studyArea,iter=100,adapt=50,burnin=50)
summary(tiger.dot$mcmc)